The Spiders (Araneae) of Bulgaria
Version: August 2018

The Spiders (Araneae) of Bulgaria

Blagoev, G., Deltshev, C., Lazarov, S. & Naumova, M.

Dr Gergin Blagoev
Biodiversity Institute of Ontario
University of Guelph
579 Gordon Street
Guelph, Ontario, N1G 2W1 CANADA

Dr Christo Deltshev
National Museum of Natural History, Bulgarian Academy of Sciences
1 Tsar Osvoboditel Blvd
1000 Sofia, BULGARIA

Dr Stoyan Lazarov
National Museum of Natural History, Bulgarian Academy of Sciences
1 Tsar Osvoboditel Blvd
1000 Sofia, BULGARIA

Dr Maria Naumova
Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences
1 Tsar Osvoboditel Blvd
1000 Sofia, BULGARIA

The present check list is based on the incorporation of all available published records on the distribution of spiders in Bulgaria. A total of 1043 spider species group taxa from 45 families were established, due to the review of 270 literature items. The principal paper is ‘A critical check list of Bulgarian spiders (Araneae)’ (Deltshev & Blagoev, 2001), where 910 species based on 173 publications, together with all taxonomic changes published in the literature are listed. Now, these data are complemented by 97 papers and 7 species are still unpublished (marked in the list with red •). This check list also contains a comprehensive list of all publications on Bulgarian spiders (in the chapter References), published between 1876 and 2018.


Historical review of arachnological studies in Bulgaria

The arachnological studies in Bulgaria started at the end of the 19th century. The first data on spiders can be found in the works of Pavessi (1876) and Hristovich (1892). However, the numerous contributions of Drensky to the study of spiders and mites are considered as the beginning of regular and continuous arachnological research. His main contributions were written before the Second World War (Drensky, 1909, 1910, 1911a, b, 1912, 1913, 1915a, b, 1921, 1924, 1925, 1926, 1927a, b, 1929, 1930, 1931a, b, c, 1932, 1933, 1934a, b, c, 1936a, b, c, 1937, 1938a, b, 1939a, b, 1940a, b, 1942a, b, 1943, 1952, 1953, 1955, 1966). His book ‘Catalogue on Spiders of the Balkan Peninsula’ is the only catalogue published so far (Drensky, 1936a). There he reported 624 spider species found on the territory of Bulgaria. After the end of the War Drensky did not continue his intensive arachnological research and his last study on spiders in the South Dobrudzha was published in 1955.

In the 10 years which follows the arachnological research almost ceased until 1967 when new data on Bulgarian araneofauna appeared as a result of the regular research started by Deltshev. Biospeleological research in Bulgaria was intensified as well. Over 600 caves were studied and 76 spider species were established of which 1 genus and 9 species newly discovered (Deltshev, 1972d, 1973b, 1974, 1975a, 1977a, b, 1978a, 1980b). Data on the spatial distribution of spiders in cave biotopes and on the coenological relationships between species was presented as well (Deltshev, 1972a, b, c, 1973b, c, 1975a, b, 1982, 1983a, 1988c). The role of spiders in some pasture ecosystems in Bulgaria and Poland was studied (Deltshev & Kayak, 1974). The ecological research was continued by Blagoev. He studied the ecology of lycosid spiders (Blagoev & Deltshev, 1989; Deltshev & Blagoev, 1994, 1995).

The research efforts of some scientists were directed towards revision of old collections and their results were published as critical surveys of different genera and families (Deltshev, 1973c, 1980b, 1983c, d, 1987a, c, 1988c, 1990, 1993; Deltshev & Blagoev; 1995; Knoflach, 1999). Some of these data was published together with Dutch and Austrian arachnologists (van Helsdingen et al., 1977; Thaler et al., 1994; van Helsdingen et al., 2001).

Comprehensive research on the araneofauna of the following Bulgarian Mountains have been carried out so far: Rila (Drensky, 1932; Deltshev, 1995b, 2000a), Pirin (Drensky, 1921; Deltshev, 1983d, 1985, 1988a, b, 1990b, 1992b; Deltshev & Blagoev, 1997), Central Balkan (Drensky, 1911a; Deltshev, 1998, 2000b; Popov et al., 2000), Vitosha (Deltshev, 1967, 1980c), Sredna Gora (Lazarov et al., 2001), Osogovo (Tzonev & Lazarov, 2001), Chepun (Dimitrov & Lazarov, 2002). The research helped the precise range differentiation of the rare, relict, endemic and endangered species with in view of their conservation and protection. Many of the data contributed to the development of the National Biodiversity Conservation Strategy and National Parks Management Plans (Deltshev et al., 1993, 1998a, 2000a, b; Popov et al., 2000a).

In recent years the development of a database of Bulgarian spiders has started. The database have been designed, organised and maintained by Blagoev (Blagoev & Georgiev, 2000a, b; Deltshev et al., 1998). The data collected was used in the preparation of ‘A critical check list of Bulgarian spiders (Araneae)’ (Deltshev & Blagoev, 2001) as well as in the newly presented Check List of Bulgarian Spiders.

Fifty eight newly discovered species group taxa in Bulgaria, described mainly by Bulgarian arachnologists, are currently regarded as valid. The first descriptions of spiders based on Bulgarian material were published by Drensky (1915). He described a total of 43 species of which 13 still regarded as valid. Rosca (1939) described one species. The rest of the newly described taxa are due to the research of: Deltshev — 25, Lazarov — 8, Dimitrov — 3, Dimitrov & Lazarov — 2, Lazarov & Naumova — 2, Bosmans — 1, van Helsdingen — 1, Deltshev & Dimitrov — 1, Deltshev & Li — 1, Logunov — 1, Azarkina & Komnenov — 1, Deltshev, Blagoev, Komnenov & Lazarov — 1 and Dimitrov, Deltshev & Lazarov — 1.

At present the group of professional Bulgarian arachnologists number six persons.


NameGeneraSpecies group taxa

Family Agelenidae

Agelena labyrinthica (Clerck, 1757)
Agelena orientalis C. L. Koch, 1841
Allagelena gracilens C. L. Koch, 1841
Coelotes atropos (Walckenaer, 1830)
Coelotes terrestris (Wider, 1834)
Eratigena agrestis (Walckenaer, 1802)
Eratigena atrica (C. L. Koch, 1843)
Eratigena picta (Simon, 1870)
Histopona breviemboli Dimitrov, Deltshev & Lazarov, 2017 [described from Bulgaria]
Histopona laeta (Kulczyński, 1897)
Histopona luxurians (Kulczyński, 1897)
Histopona torpida (C. L. Koch, 1837)
Histopona tranteevi Deltshev, 1978 [described from Bulgaria]
Inermocoelotes brevispinus (Deltshev & Dimitrov, 1996) [described from Bulgaria]
Inermocoelotes deltshevi (Dimitrov, 1996) [described from Bulgaria]
Inermocoelotes drenskii (Deltshev, 1990) [described from Bulgaria]
Inermocoelotes falciger (Kulczyński, 1897)
Inermocoelotes inermis (L. Koch, 1855)
Inermocoelotes jurinitschi (Drensky, 1915) [described from Bulgaria]
Inermocoelotes karlinskii (Kulczyński, 1906)
Inermocoelotes kulczynskii (Drensky, 1915) [described from Bulgaria]
Inermocoelotes microlepidus (de Blauwe, 1973)
Inermocoelotes xinpingwangi Deltshev, 2009 [described from Bulgaria]
Lycosoides coarctata (Dufour, 1831)
Lycosoides flavomaculata Lucas, 1846
Maimuna vestita (C. L. Koch, 1841)
Tegenaria argaeica Nosek, 1905
Tegenaria bozhkovi Deltshev, 2008 [described from Bulgaria]
Tegenaria campestris C. L. Koch, 1834
Tegenaria dalmatica Kulczyński, 1906
Tegenaria domestica (Clerck, 1757)
Tegenaria ferruginea (Panzer, 1804)
Tegenaria hasperi Chyzer, 1897
Tegenaria montana Deltshev, 1993 [described from Bulgaria]
Tegenaria pagana C. L. Koch, 1840
Tegenaria parietina (Fourcroy, 1785)
Tegenaria percuriosa Brignoli, 1972
Tegenaria regispyrrhi Brignoli, 1976
Tegenaria rilaensis Deltshev, 1993 [described from Bulgaria]
Tegenaria silvestris L. Koch, 1872
Textrix caudata L. Koch, 1872
Textrix chyzeri de Blauwe, 1980
Textrix denticulata (Olivier, 1789)
Urocoras longispinus (Kulczyński, 1897)

Family Amaurobiidae

Amaurobius deelemanae Thaler & Knoflach, 1995
Amaurobius erberi (Keyserling, 1863)
Amaurobius fenestralis (Ström, 1768)
Amaurobius ferox (Walckenaer, 1830)
Amaurobius obustus L. Koch, 1868
Amaurobius pallidus L. Koch, 1868
Amaurobius similis Blackwall, (1861)
Amaurobius strandi Charitonov, 1937
Callobius balcanicus (Drensky, 1940) [described from Bulgaria]
Callobius claustrarius (Hahn, 1833)

Family Anapidae

Zangherella relicta (Kratochvíl, 1935)

Family Anyphaenidae

Anyphaena accentuata (Walckenaer, 1802)
Anyphaena sabina L. Koch, 1866

Family Araneidae

Aculepeira armida (Audouin, 1826)
Aculepeira ceropegia (Walckenaer, 1802)
Aculepeira talishia (Zawadsky, 1902)
Agalenatea redii (Scopoli, 1763)
Araneus alsine (Walckenaer, 1802)
Araneus angulatus Clerck, 1757
Araneus circe (Audouin, 1826)
Araneus diadematus Clerck, 1757
Araneus grossus (C. L. Koch, 1844)
Araneus marmoreus Clerck, 1757
Araneus nordmanni (Thorell, 1870)
Araneus quadratus Clerck, 1757
Araneus saevus (L. Koch, 1872)
Araneus sturmi (Hahn, 1831)
Araneus triguttatus (Fabricius, 1793)
Araniella alpica (L. Koch, 1869)
Araniella cucurbitina (Clerck, 1757)
Araniella displicata (Hentz, 1847)
Araniella inconspicua (Simon, 1874)
Araniella opisthographa (Kulczyński, 1905)
Araniella proxima (Kulczyński, 1885)
Argiope bruennichi (Scopoli, 1772)
Argiope lobata (Pallas, 1772)
Cercidia prominens (Westring, 1851)
Cyclosa algerica Simon, 1885
Cyclosa conica (Pallas, 1772)
Cyclosa oculata (Walckenaer, 1802)
Cyclosa sierrae Simon, 1870
Gibbaranea bituberculata (Walckenaer, 1802)
Gibbaranea gibbosa (Walckenaer, 1802)
Gibbaranea omoeda (Thorell, 1870)
Gibbaranea ullrichi (Hahn, 1835)
Glyptogona sextuberculata (Keyserling, 1863)
Hypsosinga albovittata (Westring, 1851)
Hypsosinga heri (Hahn, 1831)
Hypsosinga pygmaea (Sundevall, 1831)
Hypsosinga sanguinea (C. L. Koch, 1844)
Larinioides cornutus (Clerck, 1757)
Larinioides ixobolus (Thorell, 1873)
Larinioides patagiatus (Clerck, 1757)
Larinioides sclopetarius (Clerck, 1757)
Larinioides suspicax O. Pickard-Cambridge, 1876
Leviellus stroemi (Thorell, 1870)
Leviellus thorelli (Ausserer, 1871)
Mangora acalypha (Walckenaer, 1802)
Neoscona adianta (Walckenaer, 1802)
Neoscona subfusca (C. L. Koch, 1837)
Nuctenea silvicultrix (C. L. Koch, 1835)
Nuctenea umbratica (Clerck, 1757)
Singa hamata (Clerck, 1757)
Singa lucina (Audouin, 1826)
Singa nitidula C. L. Koch, 1844
Singa semiatra L. Koch, 1867
Zilla diodia (Walckenaer, 1802)
Zygiella atrica (C. L. Koch, 1845)
Zygiella keyserlingi (Ausserer, 1871)
Zygiella montana (C. L. Koch, 1834)
Zygiella x-notata (Clerck, 1757)

Family Atypidae

Atypus affinis Eichwald, 1830
Atypus muralis Bertkau, 1890
Atypus piceus (Sulzer, 1776)

Family Clubionidae

Clubiona alpicola Kulczyński, 1882
Clubiona brevipes Blackwall, 1841
Clubiona caerulescens L. Koch, 1867
Clubiona comta C. L. Koch, 1839
Clubiona corticalis (Walckenaer, 1802)
Clubiona diversa O. Pickard-Cambridge, 1862
Clubiona frisia Wunderlich & Schuett, 1995
Clubiona frutetorum L. Koch, 1867
Clubiona genevensis L. Koch, 1866
Clubiona germanica Thorell, 1871
Clubiona juvenis Simon, 1878
Clubiona leucaspis Simon, 1932
Clubiona lutescens Westring, 1851
Clubiona marmorata L. Koch, 1866
Clubiona neglecta O. Pickard-Cambridge, 1862
Clubiona pallidula (Clerck, 1757)
Clubiona phragmitis C. L. Koch, 1843
Clubiona prope subsultans Thorell, 1875
Clubiona pseudoneglecta Wunderlich, 1994
Clubiona reclusa O. Pickard-Cambridge, 1863
Clubiona saxatilis L. Koch, 1866
Clubiona similis L. Koch, 1867
Clubiona subsultans Thorell, 1875
Clubiona subtilis L. Koch, 1867
Clubiona terrestris Westring, 1851
Clubiona trivialis C. L. Koch, 1843
Clubiona vegeta L. Koch, 1874

Family Cybaeidae

Cryphoeca pirini (Drensky, 1921) [described from Bulgaria]
Cryphoeca silvicola (C. L. Koch, 1834)
Cybaeus angustiarum L. Koch, 1868
Cybaeus balkanus Deltshev, 1997 [described from Bulgaria]
Tuberta maerens (O. Pickard-Cambridge, 1863)

Family Dictynidae

Archaeodictyna ammophila (Menge, 1871)
Argenna patula (Simon, 1874)
Argenna subnigra (O. Pickard-Cambridge, 1861)
Argyroneta aquatica (Clerck, 1757)
Brigittea civica (Lucas, 1850)
Brigittea latens (Fabricius, 1775)
Brigittea vicina Simon, 1873
Brommella falcigera (Balogh, 1935)
Dictyna arundinacea (Linnaeus, 1758)
Dictyna major Menge, 1869
Dictyna pusilla Thorell, 1856
Dictyna uncinata Thorell, 1856
Emblyna brevidens (Kulczyński, 1897)
Lathys humilis Blackwall, 1855
Lathys stigmatisata (Menge, 1869)
Marilynia bicolor (Simon, 1870)
Nigma flavescens (Walckenaer, 1830)
Nigma puella (Simon, 1870)
Nigma walckenaeri (Roewer, 1951)
Scotolathys simplex Simon, 1884

Family Dysderidae

Dasumia amoena (Kulczyński, 1897)
Dasumia canestrinii (L. Koch, 1876)
Dasumia kusceri (Kratochvíl, 1935)
Dysdera adriatica Kulczyński, 1897
Dysdera crocata C. L. Koch, 1838
Dysdera enguriensis Deeleman-Reinhold, 1988
Dysdera erythrina (Walckenaer, 1802)
Dysdera hungarica Kulczyński, 1897
Dysdera kollari Doblika, 1851
Dysdera lata Reuss, 1834
Dysdera longirostris Doblika, 1853
Dysdera ninnii Canestrini, 1868
Dysdera pectinata Deeleman-Reinhold, 1988
Dysdera punctata C. L. Koch, 1838
Dysdera richteri Charitonov, 1956
Dysdera westringi O. Pickard-Cambridge, 1872
Dysderocrates egregius (Kulczyński, 1897)
Harpactea abantia (Simon, 1884)
Harpactea alexandrae Lazarov, 2006 [described from Bulgaria]
Harpactea apollinea Brignoli, 1979
Harpactea asparuhi Lazarov, 2008 [described from Bulgaria]
Harpactea babori (Nosek, 1905)
Harpactea bulgarica Lazarov & Naumova, 2010 [described from Bulgaria]
Harpactea deltshevi Dimitrov & Lazarov, 1999 [described from Bulgaria]
Harpactea doblikae (Thorell, 1875)
Harpactea hombergi (Scopoli, 1763)
Harpactea konradi Lazarov, 2009 [described from Bulgaria]
Harpactea krumi Lazarov, 2010 [described from Bulgaria]
Harpactea kubrati Lazarov, 2008 [described from Bulgaria]
Harpactea lazarovi Deltshev, 2011 [described from Bulgaria]
Harpactea lepida (C. L. Koch, 1838)
Harpactea mentor Lazarov & Naumova, 2010 [described from Bulgaria]
Harpactea rubicunda (C. L. Koch, 1838)
Harpactea saeva (Herman, 1879)
Harpactea samuili Lazarov, 2006 [described from Bulgaria]
Harpactea simovi Deltshev & Lazarov, 2018 [described from Bulgaria]
Harpactea srednagora Dimitrov & Lazarov, 1999 [described from Bulgaria]
Harpactea stoevi Deltshev & Lazarov, 2018 [described from Bulgaria]
Harpactea strandjica Dimitrov, 1997 [described from Bulgaria]
Harpactea sturanyi (Nosek, 1905)
Harpactea terveli Lazarov, 2009 [described from Bulgaria]

Family Eresidae

Eresus kollari Rossi, 1846
Eresus walckenaeri Brullé, 1832

Family Eutichuridae

Cheiracanthium elegans Thorell, 1875
Cheiracanthium erraticum (Walckenaer, 1802)
Cheiracanthium macedonicum Drensky, 1921 [described from Bulgaria]
Cheiracanthium margaritae Sterghiu, 1985
Cheiracanthium mildei L. Koch, 1864
Cheiracanthium montanum L. Koch, 1877
Cheiracanthium oncognathum Thorell, 1871
Cheiracanthium pelasgicum (C. L. Koch, 1837)
Cheiracanthium pennyi O. Pickard-Cambridge, 1873
Cheiracanthium punctorium (Villers, 1789)
Cheiracanthium seidlitzi L. Koch, 1864
Cheiracanthium virescens (Sundevall, 1833)

Family Filistatidae

Filistata insidiatrix (Forskål, 1775)
Pritha nana (Simon, 1868)

Family Gnaphosidae

Aphantaulax cincta (L. Koch, 1866)
Aphantaulax trifasciata (O. Pickard-Cambridge, 1872)
Berlandina cinerea (Menge, 1872)
Berlandina nubivaga (Simon, 1878)
Callilepis concolor Simon, 1914
Callilepis cretica (Roewer, 1928)
Callilepis nocturna (Linnaeus, 1758)
Callilepis schuszteri (Herman, 1879)
Civizelotes caucasius (L. Koch, 1866)
Civizelotes gracilis (Canestrini, 1868)
Civizelotes solstitialis (Levy, 1998)
Drassodes cupreus (Blackwall, 1834)
Drassodes lapidosus (Walckenaer, 1802)
Drassodes lutescens (C. L. Koch, 1839)
Drassodes pubescens (Thorell, 1856)
Drassodes villosus (Thorell, 1856)
Drassyllus lutetianus (L. Koch, 1866)
Drassyllus praeficus (L. Koch, 1866)
Drassyllus pumilus (C. L. Koch, 1839)
Drassyllus pusillus (C. L. Koch, 1833)
Drassyllus villicus (Thorell, 1875)
Echemus angustifrons (Westring, 1861)
Gnaphosa badia (L. Koch, 1866)
Gnaphosa bicolor (Hahn, 1833)
Gnaphosa dolanskyi Řezáč, Růžička, Oger & Řezáčová, 2018 [described from Bulgaria]
Gnaphosa dolosa Herman, 1879
Gnaphosa leporina (L. Koch, 1866)
Gnaphosa lucifuga (Walckenaer, 1802)
Gnaphosa lugubris (C. L. Koch, 1839)
Gnaphosa microps Holm, 1939
Gnaphosa moesta Thorell, 1875
Gnaphosa montana (L. Koch, 1866)
Gnaphosa muscorum (L. Koch, 1866)
Gnaphosa opaca Herman, 1879
Gnaphosa rhenana Müller & Schenkel, 1895
Gnaphosa taurica Thorell, 1875
Haplodrassus aenus Thaler, 1984
Haplodrassus bohemicus Miller & Buchar, 1977
Haplodrassus cognatus (Westring, 1861)
Haplodrassus dalmatensis (L. Koch, 1866)
Haplodrassus kulczynskii Lohmander, 1942
Haplodrassus minor (O. Pickard-Cambridge, 1879)
Haplodrassus orientalis (L. Koch, 1866)
Haplodrassus signifer (C. L. Koch, 1839)
Haplodrassus silvestris (Blackwall, 1833)
Haplodrassus umbratilis (L. Koch, 1866)
Kishidaia conspicua (L. Koch, 1866)
Leptodrassus femineus (Simon, 1873)
Micaria aenea Thorell, 1871
Micaria albovittata (Lucas, 1846)
Micaria coarctata (Lucas, 1846)
Micaria dives (Lucas, 1846)
Micaria formicaria (Sundevall, 1831)
Micaria fulgens (Walckenaer, 1802)
Micaria funerea Simon, 1878
Micaria guttulata (C. L. Koch, 1839)
Micaria pallipes (Lucas, 1846)
Micaria pulicaria (Sundevall, 1831)
Micaria rossica Thorell, 1875
Micaria silesiaca L. Koch, 1875
Micaria sociabilis Kulczyński, 1897
Micaria subopaca Westring, 1861
Nomisia aussereri (L. Koch, 1872)
Nomisia exornata (C. L. Koch, 1839)
Nomisia ripariensis (O. Pickard-Cambridge, 1872)
Phaeocedus braccatus (L. Koch, 1866)
Poecilochroa variana (C. L. Koch, 1839)
Scotophaeus blackwalli (Thorell, 1871)
Scotophaeus quadripunctatus (Linnaeus, 1758)
Scotophaeus scutulatus (L. Koch, 1866)
Setaphis carmeli (O. Pickard-Cambridge, 1872)
Sosticus loricatus (L. Koch, 1866)
Trachyzelotes barbatus (L. Koch, 1866)
Trachyzelotes kulczynskii (Bösenberg, 1902)
Trachyzelotes malkini Platnick & Murphy, 1984
Trachyzelotes pedestris (C. L. Koch, 1837)
Urozelotes rusticus (L. Koch, 1872)
Zelotes aeneus (Simon, 1878)
Zelotes apricorum (L. Koch, 1876)
Zelotes atrocaeruleus (Simon, 1878)
Zelotes aurantiacus Miller, 1967
Zelotes balcanicus Deltshev, 2006 [described from Bulgaria]
Zelotes callidus (Simon, 1878)
Zelotes cingarus (O. Pickard-Cambridge, 1874)
Zelotes clivicola (L. Koch, 1870)
Zelotes electus (C. L. Koch, 1839)
Zelotes erebeus (Thorell, 1871)
Zelotes exiguus (Müller & Schenkel, 1895)
Zelotes gallicus Simon, 1914
Zelotes harmeron Levy, 2009
Zelotes hermani (Chyzer, 1897)
Zelotes latreillei (Simon, 1878)
Zelotes longipes (L. Koch, 1866)
Zelotes oblongus (C. L. Koch, 1833)
Zelotes olympi (Kulczyński, 1903)
Zelotes petrensis (C. L. Koch, 1839)
Zelotes prope olympi (Kulczyński, 1903)
Zelotes similis (Kulczyński, 1887)
Zelotes strandi (Nosek, 1905)
Zelotes subterraneus (C. L. Koch, 1833)
Zelotes talpinus (L. Koch, 1872)
Zelotes tenuis (L. Koch, 1866)
Zelotes vespertinus (Thorell, 1875)
Zelotes zellensis Grimm, 1982

Family Hahniidae

Antistea elegans (Blackwall, 1841)
Cicurina cicur (Fabricius, 1793)
Hahnia nava (Blackwall, 1841)
Hahnia ononidum Simon, 1875
Hahnia pusilla C. L. Koch, 1841
Hahniharmia picta (Kulczyński, 1897)
Iberina candida (Simon, 1875)
Mastigusa macrophthalma (Kulczyński, 1897)

Family Leptonetidae

Cataleptoneta detriticola Deltshev & Li, 2013 [described from Bulgaria]
Protoleptoneta beroni Deltshev, 1977 [described from Bulgaria]
Protoleptoneta bulgarica Deltshev, 1972 [described from Bulgaria]

Family Linyphiidae

Abacoproeces saltuum (L. Koch, 1872)
Acartauchenius scurrilis (O. Pickard-Cambridge, 1872)
Agyneta cauta (O. Pickard-Cambridge, 1902)
Agyneta equestris (L. Koch, 1881)
Agyneta fuscipalpa (C. L. Koch, 1836)
Agyneta gulosa (L. Koch, 1869)
Agyneta ramosa Jackson, 1912
Agyneta rurestris (C. L. Koch, 1836)
Agyneta simplicitarsis (Simon, 1884)
Anguliphantes angulipalpis (Westring, 1851)
Antrohyphantes balcanicus (Drensky, 1931) [described from Bulgaria]
Antrohyphantes rhodopensis (Drensky, 1931) [described from Bulgaria]
Antrohyphantes sophianus (Drensky, 1931) [described from Bulgaria]
Araeoncus anguineus (L. Koch, 1869)
Araeoncus clivifrons Deltshev, 1987 [described from Bulgaria]
Araeoncus crassiceps (Westring, 1861)
Araeoncus humilis (Blackwall, 1841)
Araeoncus tauricus Gnelitsa, 2005
Archaraeoncus prospiciens (Thorell, 1875)
Bathyphantes gracilis (Blackwall, 1841)
Bathyphantes nigrinus (Westring, 1851)
Bolyphantes alticeps (Sundevall, 1833)
Bolyphantes kolosvaryi (Caporiacco, 1936)
Bolyphantes luteolus (Blackwall, 1833)
Caviphantes dobrogicus (Dumitrescu & Miller, 1962)
Centromerita bicolor (Blackwall, 1833)
Centromerus acutidentatus Deltshev, 2002 [described from Bulgaria]
Centromerus brevipalpus (Menge, 1866)
Centromerus bulgarianus (Drensky, 1931) [described from Bulgaria]
Centromerus capucinus (Simon, 1884)
Centromerus cavernarum (L. Koch, 1872)
Centromerus incilium (L. Koch, 1881)
Centromerus lakatnikensis (Drensky, 1931) [described from Bulgaria]
Centromerus milleri Deltshev, 1974 [described from Bulgaria]
Centromerus pabulator (O. Pickard-Cambridge, 1875)
Centromerus prudens (O. Pickard-Cambridge, 1873)
Centromerus semiater (L. Koch, 1879)
Centromerus serratus (O. Pickard-Cambridge, 1875)
Centromerus silvicola (Kulczyński, 1887)
Centromerus sylvaticus (Blackwall, 1841)
Centromerus sylvaticus paucidentatus Deltshev, 1983 [described from Bulgaria]
Centromerus valkanovi Deltshev, 1983 [described from Bulgaria]
Ceratinella brevipes (Westring, 1851)
Ceratinella brevis (Wider, 1834)
Ceratinella major Kulczyński, 1894
Ceratinella scabrosa (O. Pickard-Cambridge, 1871)
Cinetata gradata (Simon, 1881)
Cresmatoneta mutinensis (Canestrini, 1868)
Crosbyarachne silvestris (Georgescu, 1973)
Dicymbium nigrum (Blackwall, 1834)
Dicymbium nigrum brevisetosum Locket, 1962
Dicymbium tibiale (Blackwall, 1836)
Diplocephalus altimontanus Deltshev, 1984 [described from Bulgaria]
Diplocephalus crassilobus (Simon, 1884)
Diplocephalus cristatus (Blackwall, 1833)
Diplocephalus graecus (O. Pickard-Cambridge, 1872)
Diplocephalus latifrons (O. Pickard-Cambridge, 1863)
Diplocephalus permixtus (O. Pickard-Cambridge, 1871)
Diplocephalus picinus (Blackwall, 1841)
Diplocephalus protuberans (O. Pickard-Cambridge, 1875)
Diplostyla concolor (Wider, 1834)
Dismodicus bifrons (Blackwall, 1841)
Dismodicus elevatus (C. L. Koch, 1838)
Donacochara speciosa (Thorell, 1875)
Drapetisca socialis (Sundevall, 1833)
Drepanotylus pirinicus Deltshev, 1992 [described from Bulgaria]
Entelecara acuminata (Wider, 1834)
Entelecara congenera (O. Pickard-Cambridge, 1879)
Entelecara flavipes (Blackwall, 1834)
Entelecara media Kulczyński, 1887
Erigone atra Blackwall, 1833
Erigone dentipalpis (Wider, 1834)
Erigone longipalpis pirini Deltshev, 1983 [described from Bulgaria]
Erigonella hiemalis (Blackwall, 1841)
Erigonoplus simplex Millidge, 1979
Erigonoplus spinifemuralis Dimitrov, 2003 [described from Bulgaria]
Evansia merens O. Pickard-Cambridge, 1900
Floronia bucculenta (Clerck, 1757)
Frontinellina frutetorum (C. L. Koch, 1834)
Gnathonarium dentatum (Wider, 1834)
Gonatium hilare (Thorell, 1875)
Gonatium nemorivagum (O. Pickard-Cambridge, 1875)
Gonatium orientale Fage, 1931
Gonatium paradoxum (L. Koch, 1869)
Gonatium rubellum (Blackwall, 1841)
Gonatium rubens (Blackwall, 1833)
Gongylidiellum latebricola (O. Pickard-Cambridge, 1871)
Gongylidiellum murcidum Simon, 1884
Gongylidium rufipes (Linnaeus, 1758)
Halorates reprobus (O. Pickard-Cambridge, 1879)
Hilaira excisa (O. Pickard-Cambridge, 1871)
Hylyphantes graminicola (Sundevall, 1830)
Hypomma cornutum (Blackwall, 1833)
Improphantes decolor (Westring, 1861)
Improphantes improbulus (Simon, 1929)
Incestophantes annulatus (Kulczyński, 1882)
Incestophantes crucifer (Menge, 1866)
Ipa keyserlingi (Ausserer, 1867)
Ipa terrenus (L. Koch, 1879)
Kratochviliella bicapitata Miller, 1938
Labulla thoracica (Wider, 1834)
Lasiargus hirsutus (Menge, 1869)
Lepthyphantes centromeroides Kulczyński, 1914
Lepthyphantes leprosus (Ohlert, 1865)
Lepthyphantes notabilis Kulczyński, 1887
Leptorhoptrum robustum (Westring, 1851)
Leptothrix hardyi (Blackwall, 1850)
Lessertia dentichelis (Simon, 1884)
Linyphia hortensis Sundevall, 1830
Linyphia triangularis (Clerck, 1757)
Macrargus carpenteri (O. Pickard-Cambridge, 1894)
Macrargus rufus (Wider, 1834)
Mansuphantes mansuetus (Thorell, 1875)
Mansuphantes prope fragilis (Thorell, 1875)
Mansuphantes rectilamellus (Deltshev, 1988) [described from Bulgaria]
Maso gallicus Simon, 1894
Maso sundevalli (Westring, 1851)
Mecopisthes peusi Wunderlich, 1972
Mecynargus paetulus (O. Pickard-Cambridge, 1875)
Megalepthyphantes collinus (L. Koch, 1872)
Megalepthyphantes nebulosus (Sundevall, 1830)
Metopobactrus orbelicus Deltshev, 1985 [described from Bulgaria]
Micrargus apertus (O. Pickard-Cambridge, 1871)
Micrargus herbigradus (Blackwall, 1854)
Micrargus subaequalis (Westring, 1851)
Microctenonyx subitaneus (O. Pickard-Cambridge, 1875)
Microlinyphia pusilla (Sundevall, 1830)
Microneta viaria (Blackwall, 1841)
Minyriolus pusillus (Wider, 1834)
Moebelia penicillata (Westring, 1851)
Mughiphantes lithoclasicola (Deltshev, 1983) [described from Bulgaria]
Mughiphantes pulcher (Kulczyński, 1881)
Nematogmus sanguinolentus (Walckenaer, 1842)
Neriene clathrata (Sundevall, 1830)
Neriene emphana (Walckenaer, 1842)
Neriene furtiva (O. Pickard-Cambridge, 1871)
Neriene montana (Clerck, 1757)
Neriene peltata (Wider, 1834)
Neriene radiata (Walckenaer, 1842)
Nusoncus nasutus Schenkel 1925
Obscuriphantes obscurus (Blackwall, 1841)
Oedothorax agrestis (Blackwall, 1853)
Oedothorax apicatus (Blackwall, 1850)
Oedothorax fuscus (Blackwall, 1834)
Oedothorax gibbifer (Kulczyński, 1882)
Oedothorax gibbosus (Blackwall, 1841)
Oedothorax retusus (Westring, 1851)
Oreoneta montigena (L. Koch, 1872)
Oreonetides glacialis (L. Koch, 1872)
Ostearius melanopygius (O. Pickard-Cambridge, 1879)
Palliduphantes alutacius (Simon, 1884)
Palliduphantes byzantinus (Fage, 1931)
Palliduphantes insignis (O. Pickard-Cambridge, 1913)
Palliduphantes istrianus (Kulczyński, 1914)
Palliduphantes pallidus (O. Pickard-Cambridge, 1871)
Palliduphantes pillichi (Kulczyński, 1915)
Palliduphantes spelaeorum (Kulczyński, 1914)
Palliduphantes trnovensis (Drensky, 1931) [described from Bulgaria]
Panamomops inconspicuus (Miller & Valesova, 1964)
Panamomops sulcifrons (Wider, 1834)
Pelecopsis elongata (Wider, 1834)
Pelecopsis mengei (Simon, 1884)
Pelecopsis parallela (Wider, 1834)
Pelecopsis radicicola (L. Koch, 1872)
Piniphantes pinicola (Simon, 1884)
Pityohyphantes phrygianus (C. L. Koch, 1836)
Pocadicnemis juncea Locket & Millidge, 1953
Pocadicnemis pumila (Blackwall, 1841)
Poeciloneta variegata (Blackwall, 1841)
Porrhomma convexum (Westring, 1851)
Porrhomma microphthalmum (O. Pickard-Cambridge, 1871)
Porrhomma microps (Roewer, 1931)
Porrhomma pygmaeum (Blackwall, 1834)
Prinerigone vagans (Audouin, 1826)
Saaristoa abnormis (Blackwall, 1841)
Sauron rayi (Simon, 1881)
Scotargus pilosus Simon, 1913
Scotinotylus alpigena (L. Koch, 1869)
Scutpelecopsis krausi (Wunderlich, 1980)
Silometopus bonessi Casemir, 1970
Silometopus reussi (Thorell, 1871)
Sintula retroversus (O. Pickard-Cambridge, 1875)
Sintula spiniger (Balogh, 1935)
Stemonyphantes lineatus (Linnaeus, 1758)
Styloctetor romanus (O. Pickard-Cambridge, 1872)
Syedra gracilis (Menge, 1869)
Tallusia experta (O. Pickard-Cambridge, 1871)
Tallusia vindobonensis (Kulczyński, 1898)
Tapinocyba biscissa (O. Pickard-Cambridge, 1872)
Tapinocyba insecta (L. Koch, 1869)
Tapinocyba mitis (O. Pickard-Cambridge, 1882)
Tapinocyba pallens (O. Pickard-Cambridge, 1872)
Tapinopa longidens (Wider, 1834)
Tenuiphantes alacris (Blackwall, 1853)
Tenuiphantes cristatus (Menge, 1866)
Tenuiphantes drenskyi (van Helsdingen, 1977) [described from Bulgaria]
Tenuiphantes flavipes (Blackwall, 1854)
Tenuiphantes floriana (van Helsdingen, 1977)
Tenuiphantes jacksoni (Schenkel, 1925)
Tenuiphantes jacksonoides (van Helsdingen, 1977)
Tenuiphantes mengei (Kulczyński, 1887)
Tenuiphantes tenebricola (Wider, 1834)
Tenuiphantes tenuis (Blackwall, 1852)
Tenuiphantes zimmermanni (Bertkau, 1890)
Theonina kratochvili Miller & Weiss, 1979
Thyreosthenius biovatus (O. Pickard-Cambridge, 1875)
Thyreosthenius parasiticus (Westring, 1851)
Tiso aestivus (L. Koch, 1872)
Tiso vagans (Blackwall, 1834)
Trichoncoides piscator (Simon, 1884)
Trichoncus affinis Kulczyński, 1894
Trichoncus auritus (L. Koch, 1869)
Trichoncus hackmani Millidge, 1956
Trichoncus saxicola (O. Pickard-Cambridge, 1861)
Trichoncyboides simoni (Lessert, 1904)
Trichopterna cito (O. Pickard-Cambridge, 1872)
Troglohyphantes bureschianus Deltshev, 1975 [described from Bulgaria]
Troglohyphantes drenskii Deltshev, 1973 [described from Bulgaria]
Troxochrus cirrifrons (O. Pickard-Cambridge, 1871)
Troxochrus scabriculus (Westring, 1851)
Typhochrestus digitatus (O. Pickard-Cambridge, 1872)
Walckenaeria acuminata Blackwall, 1833
Walckenaeria alticeps (Denis, 1952)
Walckenaeria antica (Wider, 1834)
Walckenaeria capito (Westring, 1861)
Walckenaeria corniculans (O. Pickard-Cambridge, 1875)
Walckenaeria cucullata (C. L. Koch, 1836)
Walckenaeria dysderoides (Wider, 1834)
Walckenaeria furcillata (Menge, 1869)
Walckenaeria kochi (O. Pickard-Cambridge, 1872)
Walckenaeria mitrata (Menge, 1868)
Walckenaeria monoceros (Wider, 1834)
Walckenaeria nudipalpis (Westring, 1851)
Walckenaeria obtusa Blackwall, 1836
Walckenaeria simplex Chyzer, 1894
Walckenaeria stylifrons (O. Pickard-Cambridge, 1875)
Walckenaeria vigilax (Blackwall, 1853)
Walckenaerianus esyunini Tanasevitch, 2004

Family Liocranidae

Agraecina lineata (Simon, 1878)
Agroeca brunnea (Blackwall, 1833)
Agroeca cuprea Menge, 1873
Agroeca lusatica (L. Koch, 1875)
Agroeca proxima (O. Pickard-Cambridge, 1871)
Apostenus fuscus Westring, 1851
Liocranoeca striata (Kulczyński, 1882)
Liocranum rupicola (Walckenaer, 1830)
Mesiotelus annulipes (Kulczyński, 1897)
Mesiotelus scopensis Drensky, 1935
Mesiotelus tenuissimus (L. Koch, 1866)
Sagana rutilans Thorell, 1875
Scotina celans (Blackwall, 1841)

Family Lycosidae

Alopecosa aculeata (Clerck, 1757)
Alopecosa albofasciata (Brullé, 1832)
Alopecosa cuneata (Clerck, 1757)
Alopecosa cursor (Hahn, 1831)
Alopecosa etrusca Lugetti & Tongiorgi, 1969
Alopecosa fabrilis (Clerck, 1757)
Alopecosa farinosa (Herman, 1879)
Alopecosa inquilina (Clerck, 1757)
Alopecosa pentheri (Nosek, 1905)
Alopecosa pinetorum (Thorell, 1856)
Alopecosa pulverulenta (Clerck, 1757)
Alopecosa schmidti (Hahn, 1835)
Alopecosa solitaria (Herman, 1879)
Alopecosa striatipes (C. L. Koch, 1839)
Alopecosa sulzeri (Pavesi, 1873)
Alopecosa taeniata (C. L. Koch, 1835)
Alopecosa taeniopus (Kulczyński, 1895)
Alopecosa trabalis (Clerck, 1757)
Arctosa cinerea (Fabricius, 1777)
Arctosa figurata (Simon, 1876)
Arctosa leopardus (Sundevall, 1833)
Arctosa lutetiana (Simon, 1876)
Arctosa maculata (Hahn, 1822)
Arctosa perita (Latreille, 1799)
Arctosa stigmosa (Thorell, 1875)
Arctosa tbilisiensis Mcheidze, 1946
Arctosa variana C. L. Koch, 1847
Aulonia albimana (Walckenaer, 1805)
Geolycosa vultuosa (C. L. Koch, 1838)
Hogna radiata (Latreille, 1817)
Lycosa praegrandis C. L. Koch, 1836
Lycosa singoriensis (Laxmann, 1770)
Pardosa agrestis (Westring, 1861)
Pardosa agricola (Thorell, 1856)
Pardosa alacris (C. L. Koch, 1833)
Pardosa albatula (Roewer, 1951)
Pardosa amentata (Clerck, 1757)
Pardosa atomaria (C. L. Koch, 1847)
Pardosa bifasciata (C. L. Koch, 1834)
Pardosa blanda (C. L. Koch, 1833)
Pardosa consimilis Nosek, 1905
Pardosa cribrata Simon, 1876
Pardosa drenskii Buchar, 1968
Pardosa ferruginea (L. Koch, 1870)
Pardosa hortensis (Thorell, 1872)
Pardosa incerta Nosek, 1905
Pardosa italica Tongiorgi, 1966
Pardosa luctinosa Simon, 1876
Pardosa lugubris (Walckenaer, 1802)
Pardosa mixta (Kulczyński, 1887)
Pardosa monticola (Clerck, 1757)
Pardosa morosa (L. Koch, 1870)
Pardosa nebulosa (Thorell, 1872)
Pardosa nigra (C. L. Koch, 1834)
Pardosa nigriceps (Thorell, 1856)
Pardosa paludicola (Clerck, 1757)
Pardosa palustris (Linnaeus, 1758)
Pardosa pontica (Thorell, 1875)
Pardosa prativaga (L. Koch, 1870)
Pardosa proxima (C. L. Koch, 1847)
Pardosa pullata (Clerck, 1757)
Pardosa riparia (C. L. Koch, 1833)
Pardosa roscai (Roewer, 1951)
Pardosa tasevi Buchar, 1968
Pardosa tatarica (Thorell, 1875)
Pardosa vittata (Keyserling, 1863)
Pirata piraticus (Clerck, 1757)
Pirata piscatorius (Clerck, 1757)
Pirata tenuitarsis Simon, 1876
Piratula hygrophila (Thorell, 1872)
Piratula insularis (Emerton, 1885)
Piratula knorri (Scopoli, 1763)
Piratula latitans (Blackwall, 1841)
Trabea paradoxa Simon, 1876
Trochosa hispanica Simon, 1870
Trochosa robusta (Simon, 1876)
Trochosa ruricola (De Geer, 1778)
Trochosa terricola Thorell, 1856
Xerolycosa miniata (C. L. Koch, 1834)
Xerolycosa nemoralis (Westring, 1861)

Family Mimetidae

Ero aphana (Walckenaer, 1802)
Ero cambridgei Kulczyński, 1911
Ero furcata (Villers, 1789)
Ero tuberculata (De Geer, 1778)
Mimetus laevigatus (Keyserling, 1863)

Family Miturgidae

Zora armillata Simon, 1878
Zora manicata Simon, 1878
Zora nemoralis (Blackwall, 1861)
Zora parallela Simon, 1878
Zora pardalis Simon, 1878
Zora silvestris Kulczyński, 1897
Zora spinimana (Sundevall, 1833)

Family Mysmenidae

Microdipoena jobi (Kraus, 1967)

Family Nemesiidae

Brachythele denieri (Simon, 1916)
Brachythele langourovi Lazarov, 2005 [described from Bulgaria]
Nemesia caementaria (Latreille, 1799)
Nemesia coheni Fuhn & Polenec, 1967

Family Nesticidae

Kryptonesticus beroni Deltshev, 1977 [described from Bulgaria]
Kryptonesticus eremita Simon, 1879
Nesticus cellulanus (Clerck, 1757)

Family Oecobiidae

Oecobius maculatus Simon, 1870
Uroctea durandi (Latreille, 1809)

Family Oonopidae

Oonops pulcher Templeton, 1835
Orchestina pavesii (Simon, 1873)
Silhouettella loricatula (Roewer, 1942)
Tapinesthis inermis (Simon, 1882)

Family Oxyopidae

Oxyopes heterophthalmus (Latreille, 1804)
Oxyopes lineatus Latreille, 1806
Oxyopes ramosus (Martini & Goeze, 1778)

Family Philodromidae

Philodromus albidus Kulczyński, 1911
Philodromus aureolus (Clerck, 1757)
Philodromus buxi Simon, 1884
Philodromus cespitum (Walckenaer, 1802)
Philodromus collinus C. L. Koch, 1835
Philodromus dispar Walckenaer, 1826
Philodromus emarginatus (Schrank, 1803)
Philodromus fuscomarginatus (De Geer, 1778)
Philodromus longipalpis Simon, 1870
Philodromus margaritatus (Clerck, 1757)
Philodromus marmoratus Kulczyński, 1891
Philodromus monitae Muster, 2010
Philodromus pinetorum Muster, 2009
Philodromus poecilus (Thorell, 1872)
Philodromus praedatus O. Pickard-Cambridge, 1871
Philodromus rufus Walckenaer, 1826
Philodromus vagulus Simon, 1875
Pulchellodromus glaucinus (Simon, 1870)
Rhysodromus fallax Sundevall, 1833
Rhysodromus histrio (Latreille, 1819)
Thanatus arenarius L. Koch, 1872
Thanatus atratus Simon, 1875
Thanatus flavidus Simon, 1875
Thanatus formicinus (Clerck, 1757)
Thanatus imbecillus L. Koch, 1878
Thanatus lineatipes Simon, 1870
Thanatus meronensis Levy, 1977
Thanatus oblongiusculus (Lucas, 1846)
Thanatus pictus L. Koch, 1881
Thanatus sabulosus (Menge, 1875)
Thanatus striatus C. L. Koch, 1845
Thanatus vulgaris Simon, 1870
Thanatus vulgaris creticus Kulczyński, 1903
Tibellus macellus Simon, 1875
Tibellus maritimus (Menge, 1875)
Tibellus oblongus (Walckenaer, 1802)

Family Pholcidae

Holocnemus pluchei (Scopoli, 1763)
Hoplopholcus forskali (Thorell, 1871)
Pholcus opilionoides (Schrank, 1781)
Pholcus phalangioides (Fuesslin, 1775)
Pholcus ponticus Thorell, 1875
Psilochorus simoni (Berland, 1911)
Spermophora senoculata (Dugès, 1836)

Family Phrurolithidae

Phrurolithus festivus (C. L. Koch, 1835)
Phrurolithus pullatus Kulczyński, 1897
Phrurolithus szilyi Herman, 1879

Family Pisauridae

Dolomedes plantarius (Clerck, 1757)
Pisaura mirabilis (Clerck, 1757)
Pisaura novicia (L. Koch, 1878)

Family Salticidae

Aelurillus deltshevi Azarkina & Komnenov, 2015 [described from Bulgaria]
Aelurillus v-insignitus (Clerck, 1757)
Asianellus festivus (C. L. Koch, 1834)
Attulus damini (Chyzer, 1891)
Attulus distinguendus (Simon, 1868)
Attulus penicillatus (Simon, 1875)
Attulus saltator (O. Pickard-Cambridge, 1868)
Ballus chalybeius (Walckenaer, 1802)
Carrhotus xanthogramma (Latreille, 1819)
Chalcoscirtus infimus (Simon, 1868)
Chalcoscirtus nigritus (Thorell, 1875)
Chalcoscirtus pseudoinfimus Ovtsharenko, 1978
Cyrba algerina (Lucas, 1846)
Dendryphantes hastatus (Clerck, 1757)
Dendryphantes rudis (Sundevall, 1833)
Euophrys frontalis (Walckenaer, 1802)
Euophrys herbigrada (Simon, 1871)
Euophrys petrensis C. L. Koch, 1837
Euophrys rufibarbis (Simon, 1868)
Evarcha arcuata (Clerck, 1757)
Evarcha falcata (Clerck, 1757)
Evarcha jucunda (Lucas, 1846)
Evarcha laetabunda (C. L. Koch, 1846)
Heliophanus aeneus (Hahn, 1832)
Heliophanus auratus C. L. Koch, 1835
Heliophanus cupreus (Walckenaer, 1802)
Heliophanus dampfi Schenkel, 1923
Heliophanus dubius C. L. Koch, 1835
Heliophanus flavipes (Hahn, 1832)
Heliophanus kochii Simon, 1868
Heliophanus lineiventris Simon, 1868
Heliophanus melinus L. Koch, 1867
Heliophanus patagiatus Thorell, 1875
Heliophanus rufithorax Simon, 1868
Heliophanus simplex Simon, 1868
Heliophanus tribulosus Simon, 1868
Leptorchestes berolinensis (C. L. Koch, 1846)
Leptorchestes mutilloides (Lucas, 1846)
Macaroeris flavicomis (Simon, 1884)
Macaroeris nidicolens (Walckenaer, 1802)
Marpissa muscosa (Clerck, 1757)
Marpissa nivoyi (Lucas, 1846)
Marpissa pomatia (Walckenaer, 1802)
Marpissa radiata (Grube, 1859)
Mendoza canestrinii (Ninni, 1868)
Menemerus falsificus Simon, 1868
Menemerus semilimbatus (Hahn, 1829)
Menemerus taeniatus (L. Koch, 1867)
Mogrus neglectus (Simon, 1868)
Myrmarachne formicaria (De Geer, 1778)
Neon levis (Simon, 1871)
Neon pictus Kulczyński, 1891
Neon rayi (Simon, 1875)
Neon reticulatus (Blackwall, 1853)
Pellenes arciger (Walckenaer, 1837)
Pellenes diagonalis (Simon, 1868)
Pellenes geniculatus (Simon, 1868)
Pellenes nigrociliatus (Simon, 1875)
Pellenes seriatus (Thorell, 1875)
Pellenes tripunctatus (Walckenaer, 1802)
Philaeus chrysops (Poda, 1761)
Phintella castriesiana (Grube, 1861)
Phlegra bresnieri (Lucas, 1846)
Phlegra fasciata (Hahn, 1826)
Pseudeuophrys erratica (Walckenaer, 1826)
Pseudeuophrys lanigera (Simon, 1871)
Pseudeuophrys obsoleta (Simon, 1868)
Pseudicius badius (Simon, 1868)
Pseudicius encarpatus (Walckenaer, 1802)
Pseudicius kulczynskii Nosek, 1905
Pseudicius picaceus (Simon, 1868)
Saitis barbipes (Simon, 1868)
Saitis graecus Kulczyński, 1905
Saitis tauricus Kulczyński, 1905
Salticus cingulatus (Panzer, 1797)
Salticus confusus Lucas, 1846
Salticus mutabilis Lucas, 1846
Salticus scenicus (Clerck, 1757)
Salticus zebraneus (C. L. Koch, 1837)
Sibianor aurocinctus (Ohlert, 1865)
Sitticus terebratus (Clerck, 1757)
Sittiflor atricapillus (Simon, 1882)
Sittiflor floricola (C. L. Koch, 1837)
Sittiflor inexpectus Logunov & Kronestedt, 1997
Sittiflor rupicola (C. L. Koch, 1837)
Sittipub pubescens (Fabricius, 1775)
Sittisax dzieduszyckii (L. Koch, 1870)
Sittisax saxicola (C. L. Koch, 1846)
Synageles dalmaticus (Keyserling, 1863)
Synageles hilarulus (C. L. Koch, 1846)
Synageles venator (Lucas, 1836)
Talavera aequipes (O. Pickard-Cambridge, 1871)
Talavera monticola (Kulczyński, 1884)
Yllenus arenarius Menge, 1868
Yllenus horvathi Chyzer, 1891

Family Scytodidae

Scytodes thoracica (Latreille, 1802)

Family Segestriidae

Segestria bavarica C. L. Koch, 1843
Segestria florentina (Rossi, 1790)
Segestria senoculata (Linnaeus, 1758)

Family Sparassidae

Micrommata ligurina (C. L. Koch, 1845)
Micrommata virescens (Clerck, 1757)
Olios argelasius (Walckenaer, 1805)

Family Synaphridae

Synaphris lehtineni Marusik, Gnelitsa & Kovblyuk, 2005

Family Tetragnathidae

Meta bourneti Simon, 1922
Meta menardi (Latreille, 1804)
Metellina mengei (Blackwall, 1870)
Metellina merianae (Scopoli, 1763)
Metellina segmentata (Clerck, 1757)
Pachygnatha clercki Sundevall, 1823
Pachygnatha clerckoides Wunderlich, 1985
Pachygnatha degeeri Sundevall, 1830
Pachygnatha listeri Sundevall, 1830
Tetragnatha dearmata Thorell, 1873
Tetragnatha extensa (Linnaeus, 1758)
Tetragnatha montana Simon, 1874
Tetragnatha nigrita Lendl, 1886
Tetragnatha nitens (Audouin, 1826)
Tetragnatha obtusa C. L. Koch, 1837
Tetragnatha pinicola L. Koch, 1870
Tetragnatha reimoseri (Rosca, 1939) [described from Bulgaria]
Tetragnatha striata L. Koch, 1862

Family Theridiidae

Anelosimus vittatus (C. L. Koch, 1836)
Asagena meridionalis Kulczyński, 1894
Asagena phalerata (Panzer, 1801)
Crustulina guttata (Wider, 1834)
Crustulina scabripes Simon, 1881
Crustulina sticta (O. Pickard-Cambridge, 1861)
Dipoena braccata (C. L. Koch, 1841)
Dipoena croatica (Chyzer, 1894)
Dipoena erythropus Simon (1881)
Dipoena melanogaster (C. L. Koch, 1837)
Dipoena nigroreticulata (Simon, 1879)
Dipoena torva (Thorell, 1875)
Enoplognatha afrodite Hippa & Oksala, 1983
Enoplognatha latimana Hippa & Oksala, 1982
Enoplognatha mandibularis (Lucas, 1846)
Enoplognatha mordax (Thorell, 1875)
Enoplognatha oelandica (Thorell, 1875)
Enoplognatha ovata (Clerck, 1757)
Enoplognatha penelope Hippa & Oksala, 1982
Enoplognatha quadripunctata Simon, 1884
Enoplognatha thoracica (Hahn, 1833)
Episinus angulatus (Blackwall, 1836)
Episinus maculipes Cavanna, 1876
Episinus truncatus Latreille, 1809
Euryopis episinoides (Walckenaer, 1847)
Euryopis flavomaculata (C. L. Koch, 1836)
Euryopis laeta (Westring, 1861)
Euryopis quinqueguttata Thorell, 1875
Euryopis saukea Levi, 1951
Euryopis sexalbomaculata (Lucas, 1846)
Heterotheridion nigrovariegatum (Simon, 1873)
Kochiura aulica (C. L. Koch, 1838)
Lasaeola convexa (Blackwall, 1870)
Lasaeola coracina (C. L. Koch, 1837)
Lasaeola prona (Menge, 1868)
Lasaeola tristis (Hahn, 1833)
Latrodectus tredecimguttatus (Rossi, 1790)
Neottiura bimaculata (Linnaeus, 1767)
Neottiura herbigrada (Simon, 1873)
Neottiura suaveolens (Simon, 1879)
Neottiura uncinata (Lucas, 1846)
Ohlertidion ohlerti (Thorell, 1870)
Paidiscura pallens (Blackwall, 1834)
Parasteatoda lunata (Clerck, 1757)
Parasteatoda simulans (Thorell, 1875)
Parasteatoda tabulata (Levi, 1980)
Parasteatoda tepidariorum (C. L. Koch, 1841)
Pholcomma gibbum (Westring, 1851)
Phycosoma inornatum (O. Pickard-Cambridge, 1861)
Phylloneta impressa (L. Koch, 1881)
Phylloneta sisyphia (Clerck, 1757)
Platnickina tincta (Walckenaer, 1802)
Robertus arundineti (O. Pickard-Cambridge, 1871)
Robertus frivaldszkyi (Chyzer, 1894)
Robertus lividus (Blackwall, 1836)
Robertus mediterraneus Eskov, 1987
Robertus neglectus (O. Pickard-Cambridge, 1871)
Robertus scoticus Jackson, 1914
Rugathodes bellicosus (Simon, 1873)
Sardinidion blackwalli O. Pickard-Cambridge, 1871
Simitidion simile (C. L. Koch, 1836)
Steatoda albomaculata (De Geer, 1778)
Steatoda bipunctata (Linnaeus, 1758)
Steatoda castanea (Clerck, 1757)
Steatoda grossa (C. L. Koch, 1838)
Steatoda paykulliana (Walckenaer, 1805)
Steatoda triangulosa (Walckenaer, 1802)
Theridion adrianopoli Drensky, 1915 [described from Bulgaria]
Theridion betteni Wiehle, 1960
Theridion boesenbergi Strand, 1904
Theridion cinereum Thorell, 1875
Theridion hemerobium Simon, 1914
Theridion melanurum Hahn, 1831
Theridion mystaceum L. Koch, 1870
Theridion petraeum L. Koch, 1872
Theridion pictum (Walckenaer, 1802)
Theridion pinastri L. Koch, 1872
Theridion varians Hahn, 1833

Family Theridiosomatidae

Theridiosoma gemmosum (L. Koch, 1877)

Family Thomisidae

Coriarachne depressa (C. L. Koch, 1837)
Cozyptila blackwalli (Simon, 1875)
Cozyptila thaleri Marusik & Kovblyuk, 2005
Diaea dorsata (Fabricius, 1777)
Diaea livens Simon, 1876
Ebrechtella tricuspidata (Fabricius, 1775)
Heriaeus graminicola (Doleschall, 1852)
Heriaeus hirtus (Latreille, 1819)
Heriaeus oblongus Simon, 1918
Heriaeus setiger (O. Pickard-Cambridge, 1872)
Heriaeus simoni Kulczyński, 1903
Misumena vatia (Clerck, 1757)
Monaeses paradoxus (Lucas, 1846)
Ozyptila atomaria (Panzer, 1801)
Ozyptila balcanica Deltshev, Blagoev, Komnenov & Lazarov, 2016 [described from Bulgaria]
Ozyptila brevipes (Hahn, 1826)
Ozyptila claveata (Walckenaer, 1837)
Ozyptila confluens (C. L. Koch, 1845)
Ozyptila conostyla Hippa, Koponen & Oksala, 1986
Ozyptila gertschi Kurata, 1944
Ozyptila lugubris (Kroneberg, 1875)
Ozyptila praticola (C. L. Koch, 1837)
Ozyptila pullata (Thorell, 1875)
Ozyptila rauda Simon, 1875
Ozyptila sanctuaria (O. Pickard-Cambridge, 1871)
Ozyptila scabricula (Westring, 1851)
Ozyptila simplex (O. Pickard-Cambridge, 1862)
Ozyptila trux (Blackwall, 1846)
Pistius truncatus (Pallas, 1772)
Runcinia grammica (C. L. Koch, 1837)
Synema globosum (Fabricius, 1775)
Synema plorator (O. Pickard-Cambridge, 1872)
Thomisus onustus Walckenaer, 1805
Tmarus piger (Walckenaer, 1802)
Tmarus piochardi (Simon, 1866)
Tmarus stellio Simon, 1875
Xysticus abditus Logunov, 2006 [described from Bulgaria]
Xysticus acerbus Thorell, 1872
Xysticus audax (Schrank, 1803)
Xysticus bifasciatus C. L. Koch, 1837
Xysticus bonneti Denis, 1938
Xysticus caperatus Simon, 1875
Xysticus cor Canestrini, 1873
Xysticus cristatus (Clerck, 1757)
Xysticus desidiosus Simon, 1875
Xysticus erraticus (Blackwall, 1834)
Xysticus gallicus Simon, 1875
Xysticus graecus C. L. Koch, 1837
Xysticus kempeleni Thorell, 1872
Xysticus kochi Thorell, 1872
Xysticus laetus Thorell, 1875
Xysticus lanio C. L. Koch, 1835
Xysticus lineatus (Westring, 1851)
Xysticus luctator L. Koch, 1870
Xysticus luctuosus (Blackwall, 1836)
Xysticus macedonicus Silhavy, 1944
Xysticus marmoratus Thorell, 1875
Xysticus ninnii Thorell, 1872
Xysticus nubilus Simon, 1875
Xysticus robustus (Hahn, 1832)
Xysticus sabulosus (Hahn, 1832)
Xysticus striatipes L. Koch, 1870
Xysticus tenebrosus Silhavy, 1944
Xysticus ulmi (Hahn, 1831)

Family Titanoecidae

Nurscia albomaculata (Lucas, 1846)
Nurscia albosignata Simon, 1874
Titanoeca incerta (Nosek, 1905)
Titanoeca quadriguttata (Hahn, 1833)
Titanoeca schineri L. Koch, 1872
Titanoeca tristis L. Koch, 1872
Titanoeca veteranica Herman, 1879

Family Trachelidae

Cetonana laticeps (Canestrini, 1868)
Metatrachelas rayi (Simon, 1878)
Paratrachelas maculatus (Thorell, 1875)

Family Uloboridae

Hyptiotes flavidus (Blackwall, 1862)
Hyptiotes paradoxus (C. L. Koch, 1834)
Uloborus plumipes Lucas, 1846
Uloborus walckenaerius Latreille, 1806

Family Zodariidae

Zodarion aculeatum Chyzer, 1897
Zodarion blagoevi Bosmans, 2009 [described from Bulgaria]
Zodarion epirense Brignoli, 1984
Zodarion frenatum Simon, 1884
Zodarion graecum (C. L. Koch, 1843)
Zodarion hauseri Brignoli, 1984
Zodarion italicum (Canestrini, 1868)
Zodarion morosum Denis, 1935
Zodarion ohridense Wunderlich, 1973
Zodarion pirini Drensky, 1921 [described from Bulgaria]
Zodarion thoni Nosek, 1905
Zodarion turcicum Wunderlich, 1980

Family Zoropsidae

Zoropsis lutea (Thorell, 1875)
Zoropsis spinimana (Dufour, 1820)


Antov A., Lazarov S., Deltshev C., Blagoev G. 2004. Spiders from the Sofia Region. A Faunistic and Zoogeographical Analysis. — In: L. Penev, J. Niemelä, D. J. Kotze et N. Chipev (Eds.), Ecology of the City of Sofia. Species and Communities in an Urban Environment. Pensoft Publishers, Sofia-Moscow, pp. 355-363.
Atanassov N. 1936. Die Höhlen ‘Maasite’ beim Dorfe Bela, Sliven Bezirk. — Mitteilungen der Bulgarischen Speläologischen Gesellschaft in Sofia, 1: 75-88. (In Bulgarian)
Atanassov N., Stefanov A. 1954. Die Höhle ‘Seeva Dupka’ bei der Ortschaft Malka Brestnitza, Kreisbezirk Teteven, in Hinblick aus das Karstgebiet zwischen den Flüssen Vit und Slatna-Panega. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 1: 234-275. (In Bulgarian)
Azarkina G. N., Komnenov M. 2015. Descriptions of two new species of Aelurillus Simon, 1884 (Araneae, Salticidae) from the Mediterranean, with the synonymization of A. steliosi Dobroruka, 2002. — ZooKeys, 516: 109-122.
Beron P. 1969. Sur les éléments boréo-alpins de la faune bulgare. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 30: 115-132.
Beron P. 1972. Essai sur la faune cavernicole de Bulgarie. III. Resultats des recherches biospeleologiques de 1966 a 1970. — International Journal of Speleology, 4: 285-349.
Beron P. 1978. Aperçu la composition, l’origine et la formation de la faune cavernicole de la Stara planina occidentale (Bulgarie). — International Journal of Speleology, 9: 197-220.
Beron P. 1994. Résultat des recherches biospéléologiques en Bulgarie de 1971 à 1994 et liste des animaux cavernicoles Bulgares. — Tranteeva, Sofia, 1: 1-137.
Beron P. 1999. Biodiversity of the high mountain terrestrial fauna in Bulgaria. — Historia naturalis bulgarica, 10: 13-33.
Beron P. 2015. Cave fauna of Bulgaria. — East-West Publishing, Sofia. 436 pp.
Blagoev G. 1999. Study of Wolf Spiders (Araneae: Lycosidae) in Shar Planina Mountain, Macedonia. — 1st Congress of Ecologists of the Republic Macedonia with International Participation. Proceedings. 20-24.09.1998. Ohrid, R. Macedonia, 1: 335-344.
Blagoev G. 2002. Check list of Macedonian spiders (Araneae). — Acta zoologica bulgarica, 54 (3): 9-36.
Blagoev G. 2005. A Contribution to the Knowledge of the Wolf Spiders (Araneae: Lycosidae) of Albania. — Acta zoologica bulgarica, 54 (3): 9-36.
Blagoev G. 2007. Fauna and zoogeography of wolf spiders (Araneae: Lycosidae) in Bulgaria. — In: Fet, V. et Popov, A. (Eds.), Biogeography and Ecology of Bulgaria. Series: Monographiae Biologicae, Vol. 82: 469-480.
Blagoev G., Deltshev C. 1989. [Biotopical distribution of wolf-spiders (Araneae, Lycosidae) in the Zemen Gorge, Southwestern Bulgaria]. — Ecologia, Sofia, 22: 73-80. (In Bulgarian)
Blagoev G., Georgiev V. 2000a. GEF Biodiversity Database for the Rila National Park. — In: M. Sakalian (Ed.), Biological Diversity of the Rila National Park. USAID, pp. 637-649.
Blagoev G., Georgiev V. 2000b. GEF Biodiversity Database for the Central Balkan National Park. — In: M. Sakalian (Ed.), Biological Diversity of the Central Balkan National Park. USAID, pp. 603-616.
Blagoev G., Lazarov S., Deltshev C. 2001. [Spiders (Araneae) in Kresna Gorge]. — In: P. Beron (Ed.), Biodiversity of Kresna Gorge (SW Bulgaria), pp. 85-102. (In Bulgarian)
Blauwe R. de 1980. Revision de la famille des Agelenidae (Araneae) habitant la region mediterraneene (3e partie). — Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 52(11): 1-28.
Bosmans R. 1997. Revision of the genus Zodarion Walckenaer, 1833, part II. Western and Central Europe, including Italy (Araneae: Zodariidae). — Bulletin of the British Arachnological Society, 10: 265-294.
Bosmans R. 2009. Revision of the genus Zodarion Walckenaer, 1833, part III. South East Europe and Turkey (Araneae: Zodariidae). — Contributions to Natural History, 12: 211-295.
Bosmans R. 2013. On the Gnaphosid and Lycosid spiders described by L. Giltay from the Balkans (Araneae : Gnaphosidae : Lycosidae). — Bulletin de la Société Royale Belge d'Entomologie, 149: 179-184.
Bosmans R., Keer J. van 1999. The genus Enoplognatha Pavesi, 1880 in the Mediterranean region (Araneae. Theridiidae). — Bulletin of the British Arachnological Society, 11: 209-241.
Bosmans R., Oger P. 2018. On two cases of male dimorphism in dwarf spiders (Araneae: Linyphiidae). — Arachnologische Mitteilungen, 55: 52-56.
Braun R. 1982. Deutung der angeblich neuen 'Deutschland'-Arten Bösenbergs und ihrer balkanischen 'Wiederfunde' (Arachnida: Araneida). — Senckenbergiana Biologica, 62: 355-384.
Breitling R., Bauer T., Schäfer M., Morano E., Barrientos J. A., Blick T. 2016. Phantom spiders 2: More notes on dubious spider species from Europe. — Arachnologische Mitteilungen/Arachnology Letters, 52: 50-77.
Buchar J. 1968. Zur Lycosidenfauna Bulgariens (Arachn., Araneae). — Věstník Československé Zoologické Společnosti v Praze, 32: 116-130.
Buchar J., Polenec A. 1974. Zur Lycosidenfauna Jugoslawiens (Araneae: Lycosidae). — Věstník Československé Zoologické Společnosti v Praze, 38: 81-85.
Buchar J., Thaler K. 2002. Über Pardosa atomaria (C.L. Koch) und undere Pardosa-Arten an Geröllufen in Sud- und Mitteleuropa (Araneae, Lycosidae). — Linzer Biologische Beiträge, 34: 445-465.
Buresch I. 1926. Untersuchungen über die Höhlenfauna Bulgariens. II. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 12: 17-56.
Buresch I. 1928. Die Hohlenfauna Bulgariens. Eine kurze Übersicht der Erforschungen und Zusammensetzung der Höhlen fauna Bulgariens und der darauf bezughabenden Literatur. — In: E. Csiki (Ed.), X Congrès International de Zoologie, tenue à Budapest 4-10 septembre 1927, 10: 1427-1437.
Buresch I. 1966. [Cave fauna]. — In: I. P. Gerassimov et Z. S. Gulubov (Eds.), Geography of Bulgaria, 1, Sofia, pp. 506-509. (In Bulgarian)
Buresch I., Arndt W. 1926. Die Glazialrelicte stellenden Tierarten Bulgariens und Mazedoniens. — Zeitschrift für Morphologie und Ökologie der Tiere, 5 (3): 381-405.
Deeleman-Reinhold C. L. 1977. Distribution patterns in European cave spiders. — Congrès International de Spéléologie 7(Sheffield, 1977): 25-34.
Deeleman-Reinhold C. L. 1985. Contribution à la connaissance des Lepthyphantes du groupe pallidus (Araneae, Linyphiidae) de Yugoslavie, Grece et Chypre. — Mémoires de Biospéologie, 12: 37-50.
Deeleman-Reinhold C. L., Deeleman P. R. 1988. Revision des Dysderinae (Araneae, Dysderidae), les espèces mediterranéennes occidentales exceptées. — Tijdschrift voor Entomologie, 131: 141-269.
Deltshev C. 1967. [On the studies of spiders (Araneae) in the Vitosha Mountain]. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 24: 51-56. (In Bulgarian)
Deltshev C. 1970. [Neue Daten über die Verbreitung der Gattung Meta (Araneae, Araneidae) in bulgarischen Höhlen]. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 32: 89-92. (In Bulgarian)
Deltshev C. 1972a. A review of spiders (Araneae) from Bulgarian caves. — In: C. Folk (Ed.), Proceedings of 5th Intennational Congress of Arachnology, Brno, pp. 99-104.
Deltshev C. 1972b. [A contribution to the study of spiders (Araneae) from the caves in Bulgaria]. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 34: 171-175. (In Bulgarian)
Deltshev C. 1972c. A contribution to the study of spiders (Araneae) from the caves in Bulgaria. II. Genus Lepthyphantes in Bulgarian caves. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 36: 137-147.
Deltshev C. 1972d. A new genus of Bulgarian cave spiders (Protoleptoneta bulgarica, n. g., n. sp., Leptonetidae). — International Journal of Speleology, 4: 275-283.
Deltshev C. 1973a. [A contribution to the study (Araneae) of spiders from the caves of Bulgaria. III. Ecological notes on spiders (Araneae) from the entrance parts of the caves]. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 38: 39-47. (In Bulgarian)
Deltshev C. 1973b. A new Troglohyphantes from Bulgarian caves (Araneae, Linyphiidae). — — International Journal of Speleology, 5: 103-109.
Deltshev C. 1973c. Redescription of Centromerus bulgarianus (Drensky, 1931) and Centromerus lakatnikensis (Drensky, 1931) (Araneae, Linyphiidae). — International Journal of Speleology, 5: 117-126.
Deltshev C. 1974. A new Centromerus from Bulgarian caves (Araneae, Linyphiidae). — International Journal of Speleology, 6: 81-86.
Deltshev C. 1975a. A new species (Troglohyphantes bureschianus n. sp., Araneae, Linyphiidae) from Bulgarian caves. — Acta zoologica bulgarica, 3: 99-104.
Deltshev C. 1975b. The genus Lepthyphantes in Bulgarian caves. — Proceedings of 6th Intennational Congress of Arachnology, Amsterdam, pp. 210-213.
Deltshev C. 1976a. [Spiders (Araneae) along the Bulgarian Black Sea coast]. — Terrestrial fauna of Bulgaria, Materials, Sofia, pp. 83-96. (In Bulgarian)
Deltshev C. 1976b. [On the spiders (Araneae) in the Loudogorie]. — Terrestrial fauna of Bulgaria, Materials, Sofia, pp. 251-260. (In Bulgarian)
Deltshev C. 1976c. [Faunistic, taxonomic, ecological and zoogeographical research on the spiders (Araneae) in Bulgaria]. — PhD thesis. Institute of Zoology, BAS, Sofia, pp. 1-23. (In Bulgarian)
Deltshev C. 1977a. A new Protoleptoneta from caves (Araneae, Leptonetidae). — Acta zoologica bulgarica, 1: 3-8.
Deltshev C. 1977b. Genus Nesticus (Nesticidae, Araneae) from Bulgarian caves. — In: V. Panoš (Ed.), Proceedings of 6th Intennational Congress of Speleology, Olomouc, 5: 73-78.
Deltshev C. 1978a. A new Histopona (Araneae, Agelenidae) from Bulgarian caves. — Acta zoologica bulgarica, 10: 57-59.
Deltshev C. 1978b. The origin, formation and zoogeography of troglobitic spiders of the Balkan Peninsula. — Symposia of the Zoological Society of London, 42: 345-351.
Deltshev C. 1980a. On the high altitude spiders (Araneae) in Bulgaria. — In: J. Gruber (Ed.), Proceedings of 8th Intennational Congress of Arachnology, Vienna, pp. 405-409.
Deltshev C. 1980b. [Spiders (Araneae) from the high altitude belt of the Vitosha Mountain]. — Acta zoologica bulgarica, 15: 78-92. (In Bulgarian)
Deltshev C. 1980c. A contribution to the taxonomical study of pallidus group of genus Lepthyphantes Menge (Araneae, Linyphiidae) in Bulgaria. — Acta zoologica bulgarica, 16: 44-56.
Deltshev C. 1981. [Rare and threatened Arachnida species (Arachnida, Solifugae, Araneae) in south-west Bulgaria]. — Regional Symposium Pr. 8-MAB, Blagoevgrad. BAS, Sofia, pp. 490-495. (In Bulgarian)
Deltshev C. 1982. [New data on the distribution of cave spiders (Araneae) in Bulgaria]. — Acta zoologica bulgarica, 19: 100-104. (In Bulgarian)
Deltshev C. 1983a. Zoogeographical review of Bulgarian cave spiders (Araneae). — In: L. Dinev (Ed.), Proceedings of the European Regional Conference of Speleology, Sofia, 1: 144-146.
Deltshev C. 1983b. A contribution to the study of sylvaticus group of genus Centromerus F. Dahl (Araneae, Linyphiidae) in Bulgaria. — Acta zoologica bulgarica, 21: 53-58.
Deltshev C. 1983c. Notes on the spiders of genus Erigone Audouin (Araneae, Erigonidae) in Bulgaria. — Acta zoologica bulgarica, 22: 71-75.
Deltshev C. 1983d. A contribution to the taxonomical and faunistic study of genus Lepthyphantes Menge (Araneae, Linyphiidae) from Pirin Mountain. — Acta zoologica bulgarica, 23: 25-32.
Deltshev C. 1984. A new Diplocephalus species from Bulgarian mountains (Arachnida, Araneae, Erigonidae). — Reichenbachia, 22: 91-93.
Deltshev C. 1985. A contribution to the study of the family Erigonidae (Araneae) from Pirin Mountain, Bulgaria, with a description of a new species (Metopobactrus orbelicus sp. n.). — Bulletin of the British Arachnological Society, 6: 359-366.
Deltshev C. 1987a. A critical review of genus Zodarion Walckenaer (Araneae, Zodariidae) in Bulgaria. — Acta zoologica bulgarica, 33: 19-25.
Deltshev C. 1987b. [Review of the spiders (Araneae) in the Bulgarian caves]. — Modern achievements of the Bulgarian Zoology, Sofia, pp. 21-24. (In Bulgarian)
Deltshev C. 1987c. A critical review of genus Araeoncus Simon in Bulgaria, with description of a new species (Araeoncus clivifrons sp. n.) (Arachnida, Araneae, Erigonidae). — Reichenbachia, 25: 97-102.
Deltshev C. 1988a. [Review of the species from the family Lycosidae (Araneae) from Pirin Mountain]. — In: B. Botev (Ed.), Fauna of southwestern Bulgaria, Sofia, 2: 170-175. (In Bulgarian)
Deltshev C. 1988b. A contribution to the study of genus Lepthyphantes Menge (Araneae, Linyphiidae) from the Pirin Mountain with a description of a new species (Lepthyphantes rectilamellus sp. n.). — Acta zoologica bulgarica, 36: 52-55.
Deltshev C. 1988c. The genus Fageiella Kratochvil and the genus Antrohyphantes Dumitresco (Araneae, Linyphiidae, Lepthyphanteae) in the caves of Balkan peninsula. — In: J. Haupt (Ed.), XI Europäisches Arachnologisches Colloquium. Technische Universität Berlin Dokumentation Kongresse und Tagungen, 38: 293-302.
Deltshev C. 1990a. A critical review of genus Coelotes Blackwall in Bulgaria with description of a new species (Coelotes drenskii sp. n., Araneae, Agelenidae). — Acta zoologica bulgarica, 40: 29-44.
Deltshev C. 1990b. The high-altitude spiders (Araneae) in the Pirin Mountains, Bulgaria. — Acta Zoologica Fennica, 190: 111-115.
Deltshev C. 1992a. A critical review of family Theridiidae (Araneae) in Bulgaria. — Acta zoologica bulgarica, 43: 13-22.
Deltshev C. 1992b. Drepanotylus pirinicus n. sp. from Pirin Mountain (Bulgaria), with comparative remarks on the other species of the genus (Arachnida, Araneae, Linyphiidae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 79: 173-176..
Deltshev C. 1993. The genus Tegenaria Latreille in Bulgaria: A critical review with description of two sibling species (Arachnida, Araneae: Agelenidae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 80: 167-174.
Deltshev C. 1995a. A review of family Agelenidae (Araneae) in Bulgaria. Taxonomic, faunistic and zoogeographical analysis. — European Journal of Entomology, 92: 581-588.
Deltshev C. 1995b. Spiders (Araneae) from the high altitude zone of Rila Mountain (Bulgaria). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 82: 217-225.
Deltshev C. 1996. The origin, formation and zoogeography of endemic spiders of Bulgaria (Araneae). — Revue suisse de Zoologie, vol. hors serie 1: 141-151.
Deltshev C. 1997a. A new species of Cybaeidae: Cyhaeus balkanus spec. nov. from the mountains of Balkan peninsula (Arachnida, Araneae). — Reichenbachia, 32 (1): 1-4.
Deltshev C. 1997b. Spiders (Araneae) from the coastal habitats of Shabla-Ezerets Lake, Bulgaria. — Acta zoologica bulgarica, 49: 58-63.
Deltshev C. 1998. Spiders from the high altitude zone of Central Stara Planina Mountain (Bulgaria) (Araneae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 85: 213-221.
Deltshev C. 1999. Faunistic and zoogeographical review of the Spiders (Araneae) of the Balkan peninsula. — The Journal of Arachnology. Kansas, 27: 255-261.
Deltshev C. 2000. The endemic spiders (Araneae) of the Balkan Peninsula. — In: P.Gajdos et S. Pekar (Eds.), Proceedings of the 18th European Colloquium of Arachnology, Stara Lesna, 1999. Ekologia (Bratislava), 19(3): 59-65.
Deltshev C. 2003a. Faunistic Diversity and Conservation Significanse of Spiders (Araneae). — In: D. Peev (Ed.), Rapid Ecological Assessment of Rila Monastery Nature Park, USAID, pp. 108-116.
Deltshev C. 2003b. A critical review of the taxa described by P. Drensky in the period 1915-1945. — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 90: 135-150.
Deltshev C. 2004a. A Zoogeographical review of Spiders (Araneae) in Balkan Peninsula. — In: Griffits et al. (Eds.), Balkan Biodiversity, pp. 193-200.
Deltshev C. 2004b. Spiders (Araneae) from Sandanski-Petrich Valley (SW Bulgaria). — Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reihe, 80 (1): 71-76.
Deltshev C. 2005. A new Hypomma species from Stara Planina Mountains, Bulgaria (Araneae, Linyphiidae). — Revue suisse de Zoologie 112 (1): 115-119.
Deltshev C. 2007. Fauna and zoogeography of spiders of the family Linyphiidae (Araneae) in Bulgaria. — In: V. Fet et A. Popov (Eds.), Biogeography and Ecology of Bulgaria. Series: Monographiae Biologicae, Vol. 82: 447-467.
Deltshev C. 2008. Two new spider species, Malthonica bozhkovi sp. nov. and Tegenaria paragamiani sp. nov. from Rhodopy Mountains (Bulgaria and Greece) (Araneae: Agelenidae). — Zootaxa, 1872: 37-44.
Deltshev C. 2011a. Faunistic Diversity of Spiders (Araneae) in Castanian forests of Belasitsa Mt. — Report. State and prospects of the Castanea sativa population in Belasitsa mountain: climate change adaptation; maintenance of biodiversity and sustainable ecosystem management, pp. 1-12.
Deltshev C. 2011b. Harpactea lazarovi sp.n. and H. tenuiemboli, two new spider species from Balkan Peninsula (Araneae, Dysderidae). — Zoosystematics and Evolution, 87 (2): 221-226.
Deltshev C. 2013. On the identity of the poorly known spider species Zelotes strandi (Araneae: Gnaphosidae). — Arachnologische Mitteilungen, 45: 4-7.
Deltshev C., Andreev S., Blagoev G., Golemansky V., Dobrev D., Milojkova G., Peneva V., Todorov M., Hubenov Z. 1993. [Invertebrates (non-Insecta) in Bulgaria (Protozoa, Nematoda, Oligochaeta, Mollusca, Crustacea, Myriapoda, Araneae, Acari)]. — In: M. Sakalian (Ed.), National biological diversity conservation strategy, 1, Sofia, pp. 149-244. (In Bulgarian)
Deltshev C., Andreev S., Blagoev G., Golemansky V., Milojkova G., Peneva V., Dobrev D., Todorov M., Hubenov Z. 1998a. Invertebrates (non-Insecta) in Bulgaria. — In: C. Meine (Ed.), Bulgaria’s biological diversity. Conservation status and needs assessment, 1, 2, Washington, pp. 109-161.
Deltshev C., Beron P., Blagoev G., Golemansky V., Najdenow V., Peneva V., Stoev P., Todorov M., Hubenov Z. 2000a. Faunistic diversity of invertebrates (non Insecta) in Rila National Park. — In: M. Sakalian (Ed.), Biological diversity of the Rila National Park, USAID, Sofia, pp. 249-284.
Deltshev C., Beron P., Blagoev G., Golemansky V., Peneva V., Stoev P., Todorov M., Hubenov Z. 2000b. Faunistic diversity of invertebrates (non Insecta) in Central Balkan National Park. — In: M. Sakalian (Ed.), Biological diversity of the Central Balkan National Park, USAID, Sofia, pp. 289-317.
Deltshev C., Blagoev G. 1992. A faunistic and zoogeographic analysis of spiders (Araneae) in Zemen gorge (Southwestern Bulgaria). — Acta zoologica bulgarica, 45: 26-35.
Deltshev C., Blagoev G. 1994. Biotopical distribution and seasonal activity of model species of the family Gnaphosidae (Araneae) in Zemen gorge (SW Bulgaria). — Arachnologische Mitteilungen, 1: 20-30.
Deltshev C., Blagoev G. 1995. A critical review of family Lycosidae (Araneae) in Bulgaria. — Revue arachnologique, 10 (10): 171-198.
Deltshev C., Blagoev G. 1997. The spiders of Pirin Mountain (Bulgaria). Taxonomic, faunistic and zoogeographical analysis (Araneae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 84: 269-286.
Deltshev C., Blagoev G. 1998. Order Araneae. — In: T. Michev (Ed.), Biodiversity of the Srebarna Biosphere Reserve. Checklist and bibliography, Sofia, pp. 68-69.
Deltshev C., Blagoev G. 2001. A critical checklist of Bulgarian spiders (Araneae). — Bulletin of the British Arachnological Society, 12: 110-138.
Deltshev C., Blagoev G., Hubenov Z. 1998. Conservation priorities on biodiversity of invertebrates (non-Insecta) in Bulgarian mountains. — Ambio, 27: 330-334.
Deltshev C., Blagoev G., Komnenov M., Lazarov S. 2016. Description of Ozyptila balcanica sp. n. from the Balkan Peninsula and its comparison with the closely related O. umbraculorum Simon, 1932 (Araneae: Thomisidae). — Acta zoologica bulgarica, 68 (4): 483-490.
Deltshev C., Bosmans R., de Spiegelaere W., Provoost L. 2006. Zelotes balcanicus sp. n., a new and widespread species from the Balkan Peninsula (Araneae, Gnaphosidae) A new Zelotes species. — Revue suisse de Zoologie, 113 (4): 711-716.
Deltshev C., Ćurčić B. 1997. Contribution to the knowledge of the group europaeus of Centromerus Dahl (Linyphiidae, Araneae) in the Balkan Peninsula. — Revue suisse de Zoologie, 104 (1): 49-55.
Deltshev C., Ćurčić B. 2002. Contribution to the study of the genus ‘europaeus’ Centromerus Dahl (Araneae, Linyphiidae) in caves of the Balkan Peninsula. — Revue suisse de Zoologie, 109 (1): 167-176.
Deltshev C., Dimitrov D. 1996. A new Coelotes (C. brevispinus sp. n.) from Bulgarian mountains (Araneae, Agelenidae). — Revue arachnologique, 11 (7): 77-79.
Deltshev C., Hubenov Z., Blagoev G., Dobrev D. 1998. [Modern methods of collecting, managing and keeping faunistic data]. — Historia naturalis bulgarica, 9: 143-154. (In Bulgarian)
Deltshev C., Kajak A. 1974. Analysis of a sheep pasture ecosystem in the Pieniny Mountains (the Carpathians). XVI. Effect of pasture management on the number and biomass of spiders (Araneae) in two climatic regions (the Pieniny and the Sredna Gora Mountains). — Ekologia polska, 22: 693-710.
Deltshev C., Lazarov S. 2018. Two new spider species, Harpactea simovi sp. n. and H. stoevi sp. n. (Araneae: Dysderidae), from the Balkan Peninsula. — Acta zoologica bulgarica, 70 (2): 3-7.
Deltshev C., Lazarov S., Blagoev G. 2005. Spiders (Araneae) from the Eastern Rhodopes (Bulgaria and Greece). — In: P. Beron et A. Popov (Eds.), Biodiversity of Bulgaria. 2. Biodiversity of Eastern Rhodopes (Bulgaria and Greece). Pensoft and National Museum of Natural History, Sofia, pp. 181-198.
Deltshev C., Lazarov S., Blagoev G., Naumova M. 2011 . Spiders (Araneae) from the Western Rhodopes Mts (Bulgaria). — P. Beron (Ed.), Biodiversity of Bulgaria 4. Biodiversity of Western Rhodopes (Bulgaria and Greece) II. Pensoft and National Museum of Natural History, Sofia, pp. 63-103.
Deltshev C., Lazarov S., Naumova M. 2011. A survey of spiders (Araneae) inhabiting the euedaphic soil stratum and the superficial underground compartment in Bulgaria. — Arachnologische Mitteilungen, 40: 33-46.
Deltshev C., Lazarov S., Petrov B. 2003. A Contribution to the Study of Spiders (Araneae) from the Caves of Bulgaria. — Acta zoologica bulgarica, 55 (2): 9-28.
Deltshev C., Li S. 2013. A new species of the genus Cataleptoneta from Belasitsa Mts Bulgaria (Araneae, Leptonetidae). — Acta Zootaxonomica Sinica, 38 (3): 514-519.
Deltshev C., Paraschi L. 1990. A contribution to the study of spiders (Araneae: Dysderidae, Salticidae, Agelenidae) in Greece, with a description of a new species (Malthonica spinipalpis Deltshev, sp. n. Agelenidae). — Biologia Gallo-hellenica, 17 (1): 3-12.
Deltshev C., Petrov B., Mitov P. 2005. [Faunistic diversity of Class Arachnida (non Acari) in Bulgaria -- present state, importance and per­spectives]. — A. Petrova, (Ed.), Current state of Bulgarian biodiversity — problems and perspectives. Bulgarian Bioplatform, Sofia, pp. 129-151. (In Bulgarian)
Dimitrov D. 1993. New and rare spiders (Araneae) to the fauna of Bulgaria. — In: Reports of the Second National Scientific Conference of Entomology, 25-27 October, Sofia, 1993, pp. 73-75.
Dimitrov D. 1994. A record of Achaearanea tabulata from the Balkan Peninsula (Araneae: Theridiidae). — Arachnologische Mitteilungen, 8: 77-79.
Dimitrov D. 1996a. A record of Zodarion turcicum Wunderlich from Europe with description of its unknown male (Arachnida, Araneae, Zodariidae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 83: 157-158.
Dimitrov D. 1996b. Coelotes deltshevi sp. n., a new spider species from Bulgaria (Arachnida, Araneae, Zodariidae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 83: 159-161.
Dimitrov D. 1997. Description of Harpactea strandjica, sp. n. from Bulgaria (Araneae, Dysderidae). — Bulletin of the British Arachnological Society, 10: 322.
Dimitrov D. 1999. The spider fauna of the Strandzha Mountain (South-East Bulgaria). I. Faunistic data and taxonomic remarks (Arachnida: Araneae). — Acta zoologica bulgarica, 51 (2/3): 15-26.
Dimitrov D. 2003. Erigonoplus spinifemuralis sp. n. (Araneae: Linyphidae: Erigoninae) -- a New Spider Species from Bulgaria. — Acta zoologica bulgarica, 55 (3): 33-35.
Dimitrov D., Deltshev C., Lazarov L. 2017. Description of Histopona breviemboli sp. n. from the Balkan Peninsula (Arachnida, Araneae, Agelenidae). — Zootaxa, 4311: 283-286.
Dimitrov D., Lazarov S. 1999. Two new species of Harpactea from Bulgaria (Araneae: Dysderidae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 86: 127-129.
Dimitrov D., Lazarov S. 2002. A Contribution to the Study of the Spiders (Areneae) in Chepun Mountain and Dragoman Swampland (NW Bulgaria). — Acta zoologica bulgarica, 54 (2): 47-53.
Drenovska O. 1941. [Excursion of the IVth course students-naturalists in the ‘Temnata Dupka’ Cave]. — Estestvoznanie, Sofia, 3: 155-157. (In Bulgarian)
Drensky P. 1909. Giftige Spinnen Bulgariens. — Priroda, Sofia, 15: 167-170. (In Bulgarian)
Drensky P. 1910. Die Gattung Tarentula (Sund.) in Bulgarien. — Horae Societatis Entomologicae. Rossicae, 39: 411-414. (In Bulgarian)
Drensky P. 1911a. Über die Spinnen-Fauna des Trojan-Balkans und seiner nördlichen Abhange bis der Stadt Lowetsch. — Entomologische Rundschau, 28: 41-43.
Drensky P. 1911b. Traces in the Bulgarian zoogeography. — Estestvoznanie, Sofia, 2: 1-15. (In Bulgarian)
Drensky P. 1912. The sexual dimorphism of spiders (Araneae). — Estestvoznanie, Sofia, 4: 1-8. (In Bulgarian)
Drensky P. 1913. Contribution à la faune d’araignée de la Bulgarie. — Sbornik na Balgarskata akademia na naukite, 2(1): 1-146. (In Bulgarian, French summary)
Drensky P. 1915a. Arachnides noveaux ou peu connus de Bulgarie. — La Revue de l’Académie Bulgare des Sciences, 12(5): 141-176. (In Bulgarian, French summary)
Drensky P. 1915b. Le venin des araignees et leur action sur l’organisme animal. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 7: 153-169. (In Bulgarian, French summary)
Drensky P. 1921. Contribution à l’étude des araignées de la Macédoine orientale et de Pirine planina. — Spisanie na Balgarskata Akademia na Naukite [Journal of the Bulgarian Academy of Sciences], 23: 1-50. (In Bulgarian, French summary)
Drensky P. 1924. Referate und Berichte. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 1: 20-22. (In Bulgarian)
Drensky P. 1925. Referate und Berichte. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 2: 36. (In Bulgarian)
Drensky P. 1926. Referate und Berichte im Jahre 1925. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 3: 27-29. (In Bulgarian)
Drensky P. 1927a. The black poisonous spider in Bulgaria. — Priroda, Sofia, 7: 126-127. (In Bulgarian)
Drensky P. 1927b. [Lathrodectus tredecim-guttatus Ros., eine giftige Spinne in Bulgarien]. — Priroda, Sofia, 8: 99-117. (In Bulgarian)
Drensky P. 1929. Spinnen (Araneae) aus Mittel und Süd-West Mazedonien. — Spisanie na Balgarskata Akademia na Naukite [Journal of the Bulgarian Academy of Sciences], 39: 1-76. (In Bulgarian)
Drensky P. 1930. Referate und Berichte im Jahre 1928-1929. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 5: 25-27. (In Bulgarian)
Drensky P. 1931a. Höhlen-Spinnen aus Bulgarien. — Spisanie na Balgarskata Akademia na Naukite [Journal of the Bulgarian Academy of Sciences], 44: 1-50. (In Bulgarian)
Drensky P. 1931b. Kleine entomologische Mitteilungen. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 6: 123-141. (In Bulgarian)
Drensky P. 1931c. [Our poisonous animals]. — Priroda, Sofia, 1: 113-115. (In Bulgarian)
Drensky P. 1932. Zur Kenntnis der Spinnenfauna Tscham-Kuria im Rilagebirge (Bulgarien). — Bulletin de l’Institut de Zoologie et Musée, Sofia, 15/16: 326-332. (In Bulgarian)
Drensky P. 1933. [Parangalitsa reserve and its animal world]. — Zeitschrift für Forstwesen, 19: 133-143. (In Bulgarian)
Drensky P. 1934a. [Fauna of Lovech and Troyan districts]. — Lovech and Lovchansko, Sofia, 6: 107-125. (In Bulgarian)
Drensky P. 1934b. Neue Methoden der Schädlingsbekämpfung. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 8: 65-70. (In Bulgarian)
Drensky P. 1934c. Referate und Mitteilungen in den Jahre 1932 u. 1933. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 8: 217-219. (In Bulgarian)
Drensky P. 1936a. Katalog der echten Spinnen (Araneae) der Balkanhalbinsel. — Sbornik na Balgarskata akademia na naukite, 32: 1-223.
Drensky P. 1936b. Studien über die Bulgarischen Spinnenfauna ihre ökologischen und biogeographischen Besonderheiten. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 17: 71-115. (In Bulgarian)
Drensky P. 1936c. Quelques notes sur la zoogéographie de la Bulgarie. — La Bulgarie. Devant le IV-e congrés des géographes et ethnographes Slaves, Sofia, pp. 59-65.
Drensky P. 1937. Die Spinnenfauna Bulgariens. I. Unterordnung Mygalomorphae, Family Ctenizidae und Atypidae. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 10: 259-280. (In Bulgarian)
Drensky P. 1938a. Die Spinnenfauna Bulgariens. II. Unterordnung Arachnomorphae, I Gruppe Tetrastica, Familien: Filistatidae, Dysderidae und Oonopidae. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 11: 81-113. (In Bulgarian)
Drensky P. 1938b. Zur Morphologic und Biologic einer neuen bulgarischen Spinnenart, Euxinella Strandi n. g. n. sp. — Festschrift Strand, Riga, 4: 569-574.
Drensky P. 1939a. Die Spinnenfauna Bulgariens. III. Unterordnung Arachnomorphae, II Gruppe Trionichia, Familien: Urocteidae, Uloboridae, Sicaridae, Pholcidae, Eresidae. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 12: 231-252. (In Bulgarian)
Drensky P. 1939b. Referate u. Mitteilungen. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 10: 163-168. (In Bulgarian)
Drensky P. 1940a. Die Spinnenfauna Bulgariens. IV. Unterordnung Arachnomorphae, II Gruppe Trionichia, Familien: Zodariidae, Dictynidae und Amaurobiidae. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 13: 169-194. (In Bulgarian)
Drensky P. 1940b. Fauna of Dobrudzha. — Prirodoznanie, Sofia, 2: 142-145. (In Bulgarian)
Drensky P. 1942a. Spinnenfauna Bulgariens. V. Unterordnung Arachnomorphae, II Gruppe Trionichia, Familie: Agelenidae. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 15: 33-60. (In Bulgarian)
Drensky P. 1942b. Über die Insekten-Fauna des Küstengebietes nördlich von Warna. — Mitteilungen der Bulgarischen Entomologischen Gesellschaft in Sofia, 12: 15-44. (In Bulgarian)
Drensky P. 1943. Die Spinnenfauna Bulgariens. VI. Unterordnung Arachnomorphae, II Gruppe Trionichia, Familie Euetrioidae. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 16: 219-254. (In Bulgarian)
Drensky P. 1952. [On the character of insect fauna in Dobrudzha forests]. — Priroda, Sofia, 1: 60-63. (In Bulgarian)
Drensky P. 1953. Die Schwarze-Giftspinne Latrodectus 13-guttatus Rossi in Bulgarien (Ökologie, medizinische Bedeutung und Bekämpfung). — Bulletin des Instituts de Médecine, Sofia, 8: 199-218. (In Bulgarian)
Drensky P. 1955. Fliegen und Arachnoideen aus den Wäldern und den Feldschutzwaldstreifen der Süd-Dobrudscha. — In: N. Stoyanov et B. Kitanov (Eds.), Collection Dobroudzha, Sofia, pp. 385-395. (In Bulgarian)
Drensky P. 1966. [Zoogeographical districts in the distribution of terrestrial fauna (in Bulgaria)]. — In: I. P. Gerassimov et Z. S. Gulubov (Eds.), Geography of Bulgaria, Sofia, 1: 500-506. (In Bulgarian)
Drensky P., Zimina R., Boev N. 1966. [General characteristics of the fauna]. — In: I. P. Gerassimov et Z. S. Gulubov (Eds.), Geography of Bulgaria, Sofia, 1: 485-491. (In Bulgarian)
Dumitresko M. 1971. Une araignée nouvelle des grottes de Bulgarie, Antrohyphantes rodopicus n. g., n. sp. (fam. Linyphiidae, sous-fam. Linyphiinae, série de genres Lepthyphanteae). — Travaux de l’Institut de Spéologie “Émile Racovitza”, 10: 167-174.
Dumitresko M., Georgesco M. 1970. Révision des représentants du genre Troglohyphantes des grottes de Roumanie. — Livredu Centenaire Émile G. Racovitza 1868- 1968, pp. 313-331.
Dumitresko M., Orghidan T. 1969. Nouvelles données obtenues dans l’étude de la faune lithoclasicole. — Travaux de l’Institut de Spéologie “Émile Racovitza”, 8: 55-71.
Fedoriak M. M. 2015. Scientific heritage of Aleksandru Roshka as а basis for retrospective analysis of araneofauna of Bukovyna. — Edited by Prof., Dr. S. S. Rudenko. Druk Art, Chernivtsi, 1-175. (In Ukrainian)
Fedoriak M., Moscaliuc L. A. 2013. The catalogue of "Alexandru Roşca" spider collection from the "Grigore Antipa" National Museum of Natural History (Bucharest). II. Mimetidae, Oxyopidae, Pholcidae, Pisauridae, Theridiidae. — Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 56 (2): 143-156.
Fedoriak M., Voloshyn V., Moscaliuc L. A. 2016. Scientific heritage of Alexandru Roşca: publications, spider collection, described species. — Arachnologische Mitteilungen/Arachnology Letters, 51: 85-91.
Flanczewska E. 1981. Remarks on Salticidae (Aranei) of Bulgaria. — Annales Zoologici, Warszawa, 36: 187-228.
Fuhn I., Niculescu-Burlacu F. 1971. Fam. Lycosidae. — Fauna Republicii Socialiste România (Arachnida), 5 (3): 1-253.
Fuhn I., Polenec A. 1967. Über die innerartliche Gliederung von Nemesia pannonica Herman (Arach., Araneae: Ctenizidae). — Senckenbergiana Biologica, 48: 295-300.
Fürst P.-A., Mulhauser G. 1991. Worldwide bibliography of Arachnids: Presentation of the entire key words system. — Arachnologia, 8: 4-15.
Gerhardt U. 1923. Weitere sexualbiologische Untersuchungen an Spinnen. — Archiv für Naturgeschichte, 89 (A, 10): 1-225.
Gerhardt U. 1942. Weitere Studien über die Biologic der Spinnen. — Archiv für Naturgeschichte, 90 (A, 5): 85-192.
Giltay L. 1932. Arachnides recueillis par M. d’Orchymont au cours de ses voyages aux Balkans et en Asie Mineure en 1929, 1930 et 1931. — Bulletin du Musée Royal d'Histoire Naturelle de Belgique, 8: 1-40.
Giltay L. 1933. Description des arachnides nouveaux recueillis par M. A. d’Orchymont aux Balkans et en Asie Mineure en 1929-31. — Acta pro Fauna et Flora Universali, Seria I, Zoologia, 1: 1-8.
Grimm U. 1985. Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). — Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg, 26: 1-318.
Guéorguiev V. 1966. Aperçu sur la faune cavernicole de la Bulgarie. — Bulletin de l’Institut de Zoologie et Musée, Sofia, 21: 157-181. (In Bulgarian)
Guéorguiev V. 1977. La faune troglobie terrestre de la péninsule Balkanique. Origine, formation et zoogéographie. — Sofia, Editions de L‘Academie Bulgare des Sciences, 182 pp.
Guéorguiev V., Beron P. 1962. Essai sur la faune cavernicole de Bulgarie. — Annales de spéléologie, 17: 285-411.
Hazelton M. 1970. Fauna from some caves in Bulgaria, one in Yugoslavia. — Transactions of the Cave Research Group of Great Britain (Ledbury), 12 (1): 33-37.
Helsdingen P. J. van 1969. A reclassification of the species of Linyphia Latreille based on the functioning of the genitalia (Araneida, Linyphiidae) I. — Zoologische Verhandelingen, 105: 1-303.
Helsdingen P. J. van 2016. Fauna Europaea: Araneae. Version 2016.1. —, accessed on February 2017.
Helsdingen P. J. van, Thaler K., Deltshev C. 1977. The tenuis group of Lepthyphantes Menge (Araneae, Linyphiidae). — Tijdschrift voor Entomologie, 120: 1-54.
Helsdingen P. J. van, Thaler K., Deltshev C. 2001. The European species of Bolyphantes with an attempt of a phylogenetic analysis (Araneae, Linyphiidae). — Memorie della Società Entomologica Italiana, Genova, 80: 3-35.
Hollá K., Šestáková A., Holecová M., Šebestová M. 2016. On the new record of the sheet-web spider Erigonoplus foveatus comb. nov. from Slovakia, with comments on Erigonoplus simplex (Araneae: Linyphiidae). — Arachnologische Mitteilungen/Arachnology Letters, 51: 80-84.
Holm Å., Kronestedt T. 1970. A taxonomic study of the wolf spiders of the Pardosa pullata-group (Araneae, Lycosidae). — Acta entomologica Bohemoslovaca, 67: 408-428.
Hristovich G. 1892. [Collection on investigation of the Bulgarian fauna]. — Sbornik za Narodni Umotvoreniya, Nauka i Knizhnina, Sofia, 8: 337-345. (In Bulgarian)
Jäger P. 1995. Spinnenaufsammlungen aus Ostösterreich mit vier Erstnachweisen für Österreich. — Arachnologische Mitteilungen, 9: 12-25.
Jurinitch S., Drensky P. 1917. Contribution a l’étude des araignées de Bulgarie. — Spisanie na Balgarskata Akademia na Naukite [Journal of the Bulgarian Academy of Sciences], 15: 109-136. (In Bulgarian)
Kalushkov P., Blagoev G., Deltshev C. 2008. Biodiversity of Epigeic Spiders in Genetically Modified (Bt) and Conventional (non-Bt) Potato Fields in Bulgaria. — Acta zoologica bulgarica, 60 (1): 61-69.
Knoflach B. 1992. Neue Robertus-Funde in den Alpen: R. mediterraneus Eskov und Robertus sp. (Arachnida, Aranei: Theridiidae). — Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 79: 161-171.
Knoflach B. 1996. Die Arten der Steatoda phalerata-Gruppe in Europa (Arachnida: Araneae, Theridiidae). — Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 69: 377-404.
Knoflach B. 1999. The comb-footed spider genera Neottiura and Coleosoma in Europe (Araneae, Theridiidae). — Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 72: 341-371.
Kolosváry G. 1938. Sulla fauna aracnologica della Jugoslavia. — Rassegna Faunistica, 5: 3-23.
Kostova R., Lazarov S., Bekchiev R., Goranov S., Simov N., Beshkov S. 2016. Soil and cave invertebrates from the village Gintsi area. — In:Speleological studies of Caves in Godech Municipality, Part 1. Association of Speleoclubs in Sofia, Sofia: pp. 39-51.
Kratochvil J., Miller F. 1938. Sur le probléme des araignées cavernicoles du genre Centromerus de la Péninsule balkanique. — Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, 11: 107-113.
Kronestedt T., Logunov D. 2003. Separation of two species standing as Sitticus zimmermanni (Simon, 1877) (Araneae, Salticidae), a pair of altitudinally segregated species. — Revue suisse de Zoologie, 110 (4): 855-873.
Langourov M., Lazarov S., Stoev P., Guéorguiev B., Deltshev C., Petrov B., Andreev S., Simov N., Bekchiev R., Antonova V., Ljubomirov T., Dedov I., Georgiev D. 2014. New and interesting records of the MSS and cave fauna of Vitosha Mt., Bulgaria. — In: A. Zhalov, I. Ivanov et I. Petrov (Eds.), Proceedings of Balkan Speleological Conference “Sofia’2014”, Sofia, Bulgaria, 28 – 30 March 2014, Caving Club “Helictite” Publisher, Sofia, pp. 66-76.
Lazarov S. 1998. A contribution to the study of the spiders (Araneae) in Sushtinska Sredna Gora Mountain, Bulgaria. — Historia naturalis bulgarica, 9: 27-34.
Lazarov S. 2002. A review of the family Dysderidae (Araneae) in Bulgaria: faunistic and zoogeographical analysis. — European Arachnology 2002. (F. Samu et Cs. Szinetár eds. Proceedings of the 20th European Colloquium of Arachnology, Szombathely 22-26 July 2002), pp. 259-265.
Lazarov S. 2004. A Contribution to the Study of Spiders (Araneae) in Macedonia. — Acta zoologica bulgarica, 56 (2) 155-166.
Lazarov S. 2005a. Spiders (Araneae) from Maquises in South-West Bulgaria. Part I. — Acta zoologica bulgarica, 57 (2): 145-152.
Lazarov S. 2005b. A new spider species from Bulgaria, Brachythele langourovi sp. n. (Araneae, Nemesiidae). — Revue suisse de Zoologie, 112 (1): 189-193.
Lazarov S. 2006a. A New Spider Species from Bulgaria, Harpactea alexandrae sp. n. (Araneae: Dysderidae). — Acta zoologica bulgarica, 58 (1): 13-16.
Lazarov S. 2006b. A new spider species, Harpactea samuili sp. n., from Bulgaria (Araneae: Dysderidae). — In: C. Deltshev et P. Stoev (Eds.), European Arachnology 2005, Acta zoologica bulgarica, Suppl. No. 1: 81-85.
Lazarov S. 2007a. Haplogyne spiders (Araneae) in Bulgaria: faunistic and zoogeographical analysis. — In: V. Fet et A. Popov (Eds.), Biogeography and Ecology of Bulgaria. Series: Monographiae Biologicae, Vol. 82: 481-492.
Lazarov S. 2007b. Spiders (Araneae) from the Maleshevska Mountain (SW Bulgaria). Part I. — Acta zoologica bulgarica, 59 (2): 133-144.
Lazarov S. 2008a. A new spider species, Harpactea asparuhi sp. nov., from Bulgaria (Araneae: Dysderidae) — Revista Ibérica de Aracnología, Vol. 15: 25-27.
Lazarov S. 2008b. A new spider species Harpactea kubrati sp. n. from Bulgaria, (Araneae, Dysderidae). — Acta zoologica bulgarica, 60 (2): 219-221.
Lazarov S. 2009a. Harpactea konradi, a new spider species from Bulgaria, (Araneae, Dysderidae). — Zootaxa, 2140: 33-37.
Lazarov S. 2009b. A new spider species from Bulgaria, Harpactea terveli sp. n. (Araneae, Dysderidae). — International Journal of Academic Research, 1 (2): 98-96.
Lazarov S. 2009c. Spiders (Araneae) from Maquis Forests in SW Bulgariа. Part II. — Acta zoologica bulgarica, 61 (1): 33-37.
Lazarov S. 2010. A new spider species Harpactea krumi sp. n. from Bulgaria (Araneae, Dysderidae). — Acta zoologica bulgarica, 62 (1): 27-31.
Lazarov S., Deltshev C., Blagoev G. 2001. Spiders (Araneae) of Sushtinska Sredna Gora Mountain, Bulgaria. — Acta zoologica bulgarica, 53 (1): 3-28.
Lazarov S., Naumova M. 2010. Two new Harpactea species from Bulgaria (Araneae: Dysderidae). — Revue Suisse de Zoologie, 117 (1): 101-110.
Lehtinen P. 1967. Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. — Annales Zoologici Fennici, 4: 199-468.
Logunov D. V. 2006. Notes on Xysticus kempeleni Thorell, 1872 and two closely related spider species (Araneae: Thomisidae). — Acta Arachnologica, 55 (1): 59-66.
Logunov D. V., Marusik Y. M. 1994. New data on the jumping spiders of the Palearctic fauna (Aranei Salticidae). — Arthropoda Selecta, 3 (1-2): 101-115.
Logunov D. V., Marusik Y. M. 2003. A revision of the genus Yllenus Simon, 1868 (Arachnida, Araneae, Salticidae). — Moscow: KMK Scientific Press Ltd., 2003, 167 pp.
Lugetti G., Tongiorgi P. 1969. Ricerche sul genere Alopecosa Simon (Araneae-Lycosidae). — Atti della Società Toscana di Scienze Naturali (B), 76: 1-100.
Marinov M. 2000. A contribute to the fauna of Srebarna Biosphere Reserve. — Acta zoologica bulgarica, 52 (2): 37-42.
Mikhailov K., Fet V. 1986. [Contribution to the spider fauna (Aranei) of Turkmenia. I. Families Anyphaenidae, Sparassidae, Zoridae, Clubionidae, Micariidae, Oxyopidae]. — Sborník Trudov Zoologicheskogo Muzeya MGU, Moscow State University, 24: 168-186. (In Russian)
Miller F. 1958. Eine neue Lepthyphantes (L. tranteevi sp. n.) aus den bulgarischen Höhlen. — Práce Brněnské Základny Československé Akademie Věd, 30: 577-583.
Mitov P. 1995. Opiliones (Arachnida) as a component of the food stuffs of some animals. — Annuaire de l`Universite de Sofia ‘St. Kliment Ohridski’, Faculte de Biologie, (1 – Zoologie), 86-87: 67-74.
Moscaliuc L. A. 2012. New faunistic records of spiders (Arachnida: Araneae) from Dobruja (Romania and Bulgaria). — Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa”, 55: 9-15.
Nankinov D. 2000. Threatened animals of Bulgaria. Natural Conservation Status and Distribution in Ecosystems. — Prof. Marin Drinov Acad. Publ. House and Pensoft Publishers, Sofia, pp. 1-146.
Nankinov D., Beshkov V., Deltshev C., Kalushkov P. 1995. Fauna of the Pirin and Slavianka Mountains. — In: I. Zagorchev (Ed.), Pirin: Geological guidebook, Sofia, pp. 9-10.
Naumova M. 2009. Contribution to the Study of the Spiders (Araneae) in Slavyanka Mountain (SW Bulgaria). — Biotechnology and Biotechnological Equipment, 23 (2): 104-108.
Naumova M. 2018. A Review of the Distribution of Genus Dolomedes Latreille, 1804 (Araneae: Pisauridae) in the Balkan Peninsula, with Some New Records from Bulgaria. — Acta zoologica bulgarica, 70 (in press)
Naumova M. V., Blagoev G., Lazarov S., Deltshev C. 2017. New faunistic data on the spider fauna of Bulgaria (Arachnida: Araneae) — Acta zoologica bulgarica, 71 (in press)
Naumova M., Blagoev G., Lazarov S., Deltshev C. 2008. Spiders (Araneae) from Lyulin Mountain (West Bulgariа). — Acta zoologica bulgarica, 60 (3): 267-276.
Naumova M., Hristovski S., Hristov G. 2016. Spiders (Arachnida: Araneae) from Prespa National Park, Albania. — Acta zoologica bulgarica, 68 (4): 503-511.
Naumova M., Lazarov S., Petrov B., Deltshev C. 2016. New faunistic data on the cave-dwelling spiders in the Balkan Peninsula (Araneae). — Ecologica Montenegrina, 7: 425-438.
Ovtsharenko V. 1979. Spiders of the families Gnaphosidae, Thomisidae, Lycosidae (Aranei) in the Great Caucasus. — Proceedings of the Zoological Institute, Academy of Sciences of the USSR, 85: 39-53.
Pavesi P. 1876. Gli Aracnidi Turchi. — Atti della Società Italiana di Scienze Naturali, 19: 1-27.
Pavlek M., Ribera C. 2017. Kryptonesticus deelemanae gen. et sp. nov. (Araneae, Nesticidae), with notes on the Mediterranean cave species. — European Journal of Taxonomy, 262: 1-27.
Petrişor A. 1999. The catalogue of “Alexandru Roşca” collection of Araneae (Arachnida) from “Grigore Antipa” National Museum of Natural History (Bucharest). I. — Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa”, Vol. 41: 65-78.
Petrov B., Lazarov S. 2000. Steatoda triangulosa (Walckenaer, 1802) feeding on a European Blind Snake. — Newsletter of the British Arachnological Society, 88: 9-10.
Popov A. 1969. La faune cavernicole dans la région du v. Gabare, distr. Vraca. — Annuel Spéléologie, Sofia, 1 (1968), 36-40: 43. (In Bulgarien)
Popov A., Deltshev C., Hubenov Z., Beschovski V., Dobrev D., Guéorguiev B. 2000. Invertebrate fauna. — In: A. Popov et T. Meshinev (Eds.), High mountain treeless zone of the Central Balkan National Park. Biological diversity and problems of its conservation, BSBCP, Sofia, pp. 339-416.
Popov V., Deltshev C., Blagoev G., Krusteva I., Deltshev D. 2000. Epigeobiont animal assemblages from two landscapes of the Bulgarian Black Sea coast: Relationship to habitat type, assemblage structure and biodiversity. II. Spiders (Araneae). — Acta zoologica bulgarica, 52 (1): 51-88.
Řezáč M., Růžička V., Oger P., Řezáčová V. 2018. European species of the Gnaphosa alpica complex (Araneae, Gnaphosidae). — Zootaxa, 4370: 289-294.
Roşca A. 1939a. Neue Spinnenarten aus der Dobrogea (Rumänien). — Zoologischer Anzeiger, 125 (3/4): 91-96.
Roşca A. 1939b. Araignées de Dobroudja. — Buletinul de Facultăţii Ştiinţe din Cernăuţi, Vol. 12: 246-328
Schwendinger P. 1990. A synopsis of the genus Atypus (Araneae, Atypidae). — Zoologica Scripta, 19: 353-366.
Simon E. 1884. Etudes arachnologiques. 16e mémoire. 23. Matériaux pour servir à la faune des Arachnides de la Grèce. — Annales de la Société Entomologique de France, (6) 4: 305-356.
Stoev P., Deltshev C., Bachvarova D., Doichinov A. 2014. Faunistic Diversity and Conservation Significance of the Cave Invertebrates in Special Protected Area Ponor (W Bulgaria). — Acta zoologica bulgarica, Suplement 5: 75-83.
Stojiċeviċ D. 1929. [Les Araignées de Serbie. Araneae Sund.]. — Glasnevin Museum de Historia Natural, Beograd, 19: 1-65.
Szita É., Samu F. 2000. Taxonomical review of Thanatus species (Philodromidae, Araneae) of Hungary. — Acta Zoologica Academiae Scientiarum Hungaricae, 46: 155-179.
Thaler K., Helsdingen P. J. van, Deltshev C. 1994. Vikariante Verbreitung im Artenkomplex von Lepthyphantes annulatus in Europa und ihre Deutung (Araneae, Linyphiidae). — Zoologischer Anzeiger, 232: 111-127.
Thaler K., Knoflach B. 1998. Two new species and new records of the genus Amaurobius (Araneae, Amaurobiidae) from Greece. — In: P. A. Selden (Ed.), Proceedings of the 17th European Colloquium of Arachnology, Edinburgh, British Arachnological Society, Burnham Beeches, pp. 107-114.
Tzonev G., Lazarov S. 2001. A Contribution to the Study of Spiders (Araneae) in Osogovo Mountain, South-West Bulgaria. — Acta zoologica bulgarica, 53 (2): 67-78.
World Spider Catalog 2017. World Spider Catalog. — Natural History Museum Bern, online at, version 18.0, accessed on February 2017.
Wunderlich J. 1977. Zur Kenntnis der Lepthyphantes nebulosus-Gruppe (Arachnida: Araneida: Linyphiidae). — Senckenbergiana Biologica, 58: 57-61.
Wunderlich J. 1979. Revision der europäische Arten der Gattung Micaria Westring, 1851, mit Anmerkungen zu den ubringen palaarktischen Arten (Arachnida: Araneida: Gnaphosidae). — Zoologische Beiträge (Neue Folge), 25 (für 1979) (2): 233–340.
Wunderlich J. 1984. Seltene und bisher unbekannte Wolfspinnen aus Mitteleuropa und Revision der Pardosa saltuaria-Gruppe (Arachnida: Araneae: Lycosidae). — Verhandlungen des naturwissenschaflichen Vereins in Hamburg, (N.F.) 27: 417-442.
Yurkevich M. 1904. [Twenty five years review of the Bulgarian Kingdom 1879-1904. Attempt at material collection]. — Zemlevedenie, Sofia, 1: 298. (In Bulgarian).


We are much obliged to all colleagues, who helped us providing references and information on the distribution of the spiders in Bulgaria. A special thank goes to Stanislav Abadjiev (lepidopterologist, NMNHS) who arranged the new version of the site. We are very grateful also to Peter van Helsdingen, Wolfgang Nentwig and collaborators and to Norman Platnick for making available to all arachnologists the websites ‘Fauna Europaea’, ‘Spiders of Europe’ and ‘The World Spider Catalog’.

The Spiders (Araneae) of Bulgaria

This check list should be cited as follows:
Blagoev, G., Deltshev, C., Lazarov, S. & Naumova, M. 2002—2018. The Spiders (Araneae) of Bulgaria. Version: August 2018. — National Museum of Natural History, Bulgarian Academy of Sciences, online at