The Spiders (Araneae) of Bulgaria
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The Spiders (Araneae) of Bulgaria

Blagoev, G., Deltshev, C. & Lazarov, S.

Dr Gergin Blagoev
Biodiversity Institute of Ontario
University of Guelph
579 Gordon Street
Guelph, Ontario, N1G 2W1 CANADA


Dr Christo Deltshev
National Museum of Natural History, Bulgarian Academy of Sciences
1 Tsar Osvoboditel Blvd
1000 Sofia, BULGARIA


Dr Stoyan Lazarov
National Museum of Natural History, Bulgarian Academy of Sciences
1 Tsar Osvoboditel Blvd
1000 Sofia, BULGARIA


The present check list is based on the incorporation of all available published records on the distribution of spiders in Bulgaria. A total of 1038 spider species group taxa from 43 families were established, due to the review of 248 literature items. The principal paper is ‘A critical check list of Bulgarian spiders (Araneae)’ (Deltshev & Blagoev, 2001), where 910 species based on 173 publications, together with all taxonomic changes published in the literature are listed. Now, these data are complemented by 75 papers and 13 species are still unpublished (marked in the list with red •). This check list also contains a comprehensive list of all publications on Bulgarian spiders (in the chapter References), published between 1876 and 2015.

Introduction


Historical review of arachnological studies in Bulgaria

The arachnological studies in Bulgaria started at the end of the 19th century. The first data on spiders can be found in the works of Pavessi (1876) and Hristovich (1892). However, the numerous contributions of Drensky to the study of spiders and mites are considered as the beginning of regular and continuous arachnological research. His main contributions were written before the Second World War (Drensky, 1909, 1910, 1911a, b, 1912, 1913, 1915a, b, 1921, 1924, 1925, 1926, 1927a, b, 1929, 1930, 1931a, b, c, 1932, 1933, 1934a, b, c, 1936a, b, c, 1937, 1938a, b, 1939a, b, 1940a, b, 1942a, b, 1943, 1952, 1953, 1955, 1966). His book ‘Catalogue on Spiders of the Balkan Peninsula’ is the only catalogue published so far (Drensky, 1936a). There he reported 624 spider species found on the territory of Bulgaria. After the end of the War Drensky did not continue his intensive arachnological research and his last study on spiders in the South Dobrudzha was published in 1955.

In the 10 years which follows the arachnological research almost ceased until 1967 when new data on Bulgarian araneofauna appeared as a result of the regular research started by Deltshev. Biospeleological research in Bulgaria was intensified as well. Over 600 caves were studied and 76 spider species were established of which 1 genus and 9 species newly discovered (Deltshev, 1972d, 1973b, 1974, 1975a, 1977a, b, 1978a, 1980b). Data on the spatial distribution of spiders in cave biotopes and on the coenological relationships between species was presented as well (Deltshev, 1972a, b, c, 1973b, c, 1975a, b, 1982, 1983a, 1988c). The role of spiders in some pasture ecosystems in Bulgaria and Poland was studied (Deltshev & Kayak, 1974). The ecological research was continued by Blagoev. He studied the ecology of lycosid spiders (Blagoev & Deltshev, 1989; Deltshev & Blagoev, 1994, 1995).

The research efforts of some scientists were directed towards revision of old collections and their results were published as critical surveys of different genera and families (Deltshev, 1973c, 1980b, 1983c, d, 1987a, c, 1988c, 1990, 1993; Deltshev & Blagoev; 1995; Knoflach, 1999). Some of these data was published together with Dutch and Austrian arachnologists (van Helsdingen et al., 1977; Thaler et al., 1994; van Helsdingen et al., 2001).

Comprehensive research on the araneofauna of the following Bulgarian Mountains have been carried out so far: Rila (Drensky, 1932; Deltshev, 1995b, 2000a), Pirin (Drensky, 1921; Deltshev, 1983d, 1985, 1988a, b, 1990b, 1992b; Deltshev & Blagoev, 1997), Central Balkan (Drensky, 1911a; Deltshev, 1998, 2000b; Popov et al., 2000), Vitosha (Deltshev, 1967, 1980c), Sredna Gora (Lazarov et al., 2001), Osogovo (Tzonev & Lazarov, 2001), Chepun (Dimitrov & Lazarov, 2002). The research helped the precise range differentiation of the rare, relict, endemic and endangered species with in view of their conservation and protection. Many of the data contributed to the development of the National Biodiversity Conservation Strategy and National Parks Management Plans (Deltshev et al., 1993, 1998a, 2000a, b; Popov et al., 2000a).

In recent years the development of a database of Bulgarian spiders has started. The database have been designed, organised and maintained by Blagoev (Blagoev & Georgiev, 2000a, b; Deltshev et al., 1998). The data collected was used in the preparation of ‘A critical check list of Bulgarian spiders (Araneae)’ (Deltshev & Blagoev, 2001) as well as in the newly presented Check List of Bulgarian Spiders.

Fifty eight newly discovered species group taxa in Bulgaria, described mainly by Bulgarian arachnologists, are currently regarded as valid. The first descriptions of spiders based on Bulgarian material were published by Drensky (1915). He described a total of 43 species of which 13 still regarded as valid. The rest of the newly described taxa are due to the research of: Deltshev — 25, Lazarov — 8, Dimitrov — 3, Dimitrov & Lazarov — 2, Lazarov & Naumova — 2, Bosmans — 1, van Helsdingen — 1, Deltshev & Dimitrov — 1, Deltshev & Li — 1 and Logunov — 1.

At present the group of professional Bulgarian arachnologists number six persons.

Families


NameGeneraSpecies group taxa
Agelenidae1147
Amaurobiidae29
Anapidae11
Anyphaenidae12
Araneidae2057
Atypidae12
Clubionidae127
Cybaeidae23
Dictynidae1019
Dysderidae440
Eresidae12
Eutichuridae113
Filistatidae22
Gnaphosidae21102
Hahniidae69
Leptonetidae23
Linyphiidae106239
Liocranidae813
Lycosidae1282
Mimetidae25
Miturgidae17
Nemesiidae24
Nesticidae13
Oecobiidae11
Oonopidae44
Oxyopidae13
Philodromidae436
Pholcidae58
Phrurolithidae13
Pisauridae23
Salticidae3295
Scytodidae11
Segestriidae13
Sparassidae23
Tetragnathidae417
Theridiidae2578
Theridiosomatidae11
Thomisidae1464
Titanoecidae27
Trachelidae33
Uloboridae24
Zodariidae111
Zoropsidae12
Total3251038

Family Agelenidae

Agelena labyrinthica (Clerck, 1757)
Agelena orientalis C. L. Koch, 1841
Allagelena gracilens C. L. Koch, 1841
Coelotes atropos (Walckenaer, 1830)
Coelotes pabulator Simon, 1875
Coelotes terrestris (Wider, 1834)
Eratigena agrestis (Walckenaer, 1802)
Eratigena atrica (C. L. Koch, 1843)
Eratigena picta (Simon, 1870)
Histopona laeta (Kulczyński, 1897)
Histopona luxurians (Kulczyński, 1897)
Histopona myops (Simon, 1885)
Histopona torpida (C. L. Koch, 1837)
Histopona tranteevi Deltshev, 1978 [described from Bulgaria]
Inermocoelotes brevispinus (Deltshev & Dimitrov, 1996) [described from Bulgaria]
Inermocoelotes deltshevi (Dimitrov, 1996) [described from Bulgaria]
Inermocoelotes drenskii (Deltshev, 1990) [described from Bulgaria]
Inermocoelotes falciger (Kulczyński, 1897)
Inermocoelotes inermis (L. Koch, 1855)
Inermocoelotes jurinitschi (Drensky, 1915) [described from Bulgaria]
Inermocoelotes karlinskii (Kulczyński, 1906)
Inermocoelotes kulczynskii (Drensky, 1915) [described from Bulgaria]
Inermocoelotes microlepidus (de Blauwe, 1973)
Inermocoelotes xinpingwangi Deltshev, 2009 [described from Bulgaria]
Lycosoides coarctata (Dufour, 1831)
Lycosoides flavomaculata Lucas, 1846
Maimuna vestita (C. L. Koch, 1841)
Tegenaria argaeica Nosek, 1905
Tegenaria bithyniae Brignoli, 1978
Tegenaria bozhkovi Deltshev, 2008 [described from Bulgaria]
Tegenaria campestris C. L. Koch, 1834
Tegenaria dalmatica Kulczyński, 1906
Tegenaria domestica (Clerck, 1757)
Tegenaria ferruginea (Panzer, 1804)
Tegenaria hasperi Chyzer, 1897
Tegenaria montana Deltshev, 1993 [described from Bulgaria]
Tegenaria pagana C. L. Koch, 1840
Tegenaria parietina (Fourcroy, 1785)
Tegenaria pasquinii Brignoli, 1978
Tegenaria percuriosa Brignoli, 1972
Tegenaria regispyrrhi Brignoli, 1976
Tegenaria rilaensis Deltshev, 1993 [described from Bulgaria]
Tegenaria silvestris L. Koch, 1872
Textrix caudata L. Koch, 1872
Textrix chyzeri de Blauwe, 1980
Textrix denticulata (Olivier, 1789)
Urocoras longispinus (Kulczyński, 1897)

Family Amaurobiidae

Amaurobius deelemanae Thaler & Knoflach, 1995
Amaurobius erberi (Keyserling, 1863)
Amaurobius fenestralis (Ström, 1768)
Amaurobius ferox (Walckenaer, 1830)
Amaurobius obustus L. Koch, 1868
Amaurobius pallidus L. Koch, 1868
Amaurobius strandi Charitonov, 1937
Callobius balcanicus (Drensky, 1940) [described from Bulgaria]
Callobius claustrarius (Hahn, 1833)

Family Anapidae

Zangherella relicta (Kratochvíl, 1935)

Family Anyphaenidae

Anyphaena accentuata (Walckenaer, 1802)
Anyphaena sabina L. Koch, 1866

Family Araneidae

Aculepeira armida (Audouin, 1826)
Aculepeira ceropegia (Walckenaer, 1802)
Aculepeira talishia (Zawadsky, 1902)
Agalenatea redii (Scopoli, 1763)
Araneus alsine (Walckenaer, 1802)
Araneus angulatus Clerck, 1757
Araneus circe (Audouin, 1826)
Araneus diadematus Clerck, 1757
Araneus grossus (C. L. Koch, 1844)
Araneus marmoreus Clerck, 1757
Araneus nordmanni (Thorell, 1870)
Araneus quadratus Clerck, 1757
Araneus saevus (L. Koch, 1872)
Araneus sturmi (Hahn, 1831)
Araneus triguttatus (Fabricius, 1793)
Araniella alpica (L. Koch, 1869)
Araniella cucurbitina (Clerck, 1757)
Araniella displicata (Hentz, 1847)
Araniella inconspicua (Simon, 1874)
Araniella opisthographa (Kulczyński, 1905)
Araniella proxima (Kulczyński, 1885)
Argiope bruennichi (Scopoli, 1772)
Argiope lobata (Pallas, 1772)
Cercidia prominens (Westring, 1851)
Cyclosa algerica Simon, 1885
Cyclosa conica (Pallas, 1772)
Cyclosa oculata (Walckenaer, 1802)
Cyclosa sierrae Simon, 1870
Gibbaranea bituberculata (Walckenaer, 1802)
Gibbaranea gibbosa (Walckenaer, 1802)
Gibbaranea omoeda (Thorell, 1870)
Gibbaranea ullrichi (Hahn, 1835)
Glyptogona sextuberculata (Keyserling, 1863)
Hypsosinga albovittata (Westring, 1851)
Hypsosinga heri (Hahn, 1831)
Hypsosinga pygmaea (Sundevall, 1831)
Hypsosinga sanguinea (C. L. Koch, 1844)
Larinioides cornutus (Clerck, 1757)
Larinioides folium (Schrank, 1803)
Larinioides ixobolus (Thorell, 1873)
Larinioides patagiatus (Clerck, 1757)
Leviellus thorelli (Ausserer, 1871)
Mangora acalypha (Walckenaer, 1802)
Neoscona adianta (Walckenaer, 1802)
Neoscona subfusca (C. L. Koch, 1837)
Nuctenea silvicultrix (C. L. Koch, 1835)
Nuctenea umbratica (Clerck, 1757)
Parazygiella montana (C. L. Koch, 1834)
Singa hamata (Clerck, 1757)
Singa lucina (Audouin, 1826)
Singa nitidula C. L. Koch, 1844
Singa semiatra L. Koch, 1867
Stroemiellus stroemi (Thorell, 1870)
Zilla diodia (Walckenaer, 1802)
Zygiella atrica (C. L. Koch, 1845)
Zygiella keyserlingi (Ausserer, 1871)
Zygiella x-notata (Clerck, 1757)

Family Atypidae

Atypus affinis Eichwald, 1830
Atypus piceus (Sulzer, 1776)

Family Clubionidae

Clubiona alpicola Kulczyński, 1882
Clubiona brevipes Blackwall, 1841
Clubiona caerulescens L. Koch, 1867
Clubiona comta C. L. Koch, 1839
Clubiona corticalis (Walckenaer, 1802)
Clubiona diversa O. Pickard-Cambridge, 1862
Clubiona frisia Wunderlich & Schuett, 1995
Clubiona frutetorum L. Koch, 1867
Clubiona genevensis L. Koch, 1866
Clubiona germanica Thorell, 1871
Clubiona juvenis Simon, 1878
Clubiona leucaspis Simon, 1932
Clubiona lutescens Westring, 1851
Clubiona marmorata L. Koch, 1866
Clubiona neglecta O. Pickard-Cambridge, 1862
Clubiona pallidula (Clerck, 1757)
Clubiona phragmitis C. L. Koch, 1843
Clubiona prope subsultans Thorell, 1875
Clubiona pseudoneglecta Wunderlich, 1994
Clubiona reclusa O. Pickard-Cambridge, 1863
Clubiona saxatilis L. Koch, 1866
Clubiona similis L. Koch, 1867
Clubiona subsultans Thorell, 1875
Clubiona subtilis L. Koch, 1867
Clubiona terrestris Westring, 1851
Clubiona trivialis C. L. Koch, 1843
Clubiona vegeta L. Koch, 1874

Family Cybaeidae

Argyroneta aquatica (Clerck, 1757)
Cybaeus angustiarum L. Koch, 1868
Cybaeus balkanus Deltshev, 1997 [described from Bulgaria]

Family Dictynidae

Archaeodictyna ammophila (Menge, 1871)
Argenna patula (Simon, 1874)
Argenna subnigra (O. Pickard-Cambridge, 1861)
Brommella falcigera (Balogh, 1935)
Cicurina cicur (Fabricius, 1793)
Dictyna arundinacea (Linnaeus, 1758)
Dictyna civica (Lucas, 1850)
Dictyna latens (Fabricius, 1775)
Dictyna pusilla Thorell, 1856
Dictyna uncinata Thorell, 1856
Dictyna vicina Simon, 1873
Lathys humilis Blackwall, 1855
Lathys stigmatisata (Menge, 1869)
Marilynia bicolor (Simon, 1870)
Mastigusa macrophthalma (Kulczyński, 1897)
Nigma flavescens (Walckenaer, 1830)
Nigma puella (Simon, 1870)
Nigma walckenaeri (Roewer, 1951)
Scotolathys simplex Simon, 1884

Family Dysderidae

Dasumia amoena (Kulczyński, 1897)
Dasumia canestrinii (L. Koch, 1876)
Dasumia kusceri (Kratochvíl, 1935)
Dysdera adriatica Kulczyński, 1897
Dysdera argaeica Nosek, 1905
Dysdera crocata C. L. Koch, 1838
Dysdera enguriensis Deeleman-Reinhold, 1988
Dysdera erythrina (Walckenaer, 1802)
Dysdera hungarica Kulczyński, 1897
Dysdera kollari Doblika, 1851
Dysdera lata Reuss, 1834
Dysdera longirostris Doblika, 1853
Dysdera ninnii Canestrini, 1868
Dysdera pectinata Deeleman-Reinhold, 1988
Dysdera punctata C. L. Koch, 1838
Dysdera richteri Charitonov, 1956
Dysdera westringi O. Pickard-Cambridge, 1872
Dysderocrates egregius (Kulczyński, 1897)
Harpactea abantia (Simon, 1884)
Harpactea alexandrae Lazarov, 2006 [described from Bulgaria]
Harpactea apollinea Brignoli, 1979
Harpactea asparuhi Lazarov, 2008 [described from Bulgaria]
Harpactea babori (Nosek, 1905)
Harpactea bulgarica Lazarov & Naumova, 2010 [described from Bulgaria]
Harpactea deltshevi Dimitrov & Lazarov, 1999 [described from Bulgaria]
Harpactea doblikae (Thorell, 1875)
Harpactea hombergi (Scopoli, 1763)
Harpactea konradi Lazarov, 2009 [described from Bulgaria]
Harpactea krumi Lazarov, 2010 [described from Bulgaria]
Harpactea kubrati Lazarov, 2008 [described from Bulgaria]
Harpactea lazarovi Deltshev, 2011 [described from Bulgaria]
Harpactea lepida (C. L. Koch, 1838)
Harpactea mentor Lazarov & Naumova, 2010 [described from Bulgaria]
Harpactea rubicunda (C. L. Koch, 1838)
Harpactea saeva (Herman, 1879)
Harpactea samuili Lazarov, 2006 [described from Bulgaria]
Harpactea srednagora Dimitrov & Lazarov, 1999 [described from Bulgaria]
Harpactea strandjica Dimitrov, 1997 [described from Bulgaria]
Harpactea sturanyi (Nosek, 1905)
Harpactea terveli Lazarov, 2009 [described from Bulgaria]

Family Eresidae

Eresus kollari Rossi, 1846
Eresus walckenaeri Brullé, 1832

Family Eutichuridae

Cheiracanthium elegans Thorell, 1875
Cheiracanthium erraticum (Walckenaer, 1802)
Cheiracanthium macedonicum Drensky, 1921 [described from Bulgaria]
Cheiracanthium margaritae Sterghiu, 1985
Cheiracanthium mildei L. Koch, 1864
Cheiracanthium montanum L. Koch, 1877
Cheiracanthium oncognathum Thorell, 1871
Cheiracanthium pelasgicum (C. L. Koch, 1837)
Cheiracanthium pennyi O. Pickard-Cambridge, 1873
Cheiracanthium punctorium (Villers, 1789)
Cheiracanthium rupestre Herman, 1879
Cheiracanthium seidlitzi L. Koch, 1864
Cheiracanthium virescens (Sundevall, 1833)

Family Filistatidae

Filistata insidiatrix (Forskål, 1775)
Pritha nana (Simon, 1868)

Family Gnaphosidae

Aphantaulax cincta (L. Koch, 1866)
Aphantaulax trifasciata (O. Pickard-Cambridge, 1872)
Berlandina cinerea (Menge, 1872)
Berlandina nubivaga (Simon, 1878)
Callilepis concolor Simon, 1914
Callilepis cretica (Roewer, 1928)
Callilepis nocturna (Linnaeus, 1758)
Callilepis schuszteri (Herman, 1879)
Civizelotes caucasius (L. Koch, 1866)
Civizelotes gallicus (Simon, 1914)
Civizelotes solstitialis (Levy, 1998)
Drassodes cupreus (Blackwall, 1834)
Drassodes lapidosus (Walckenaer, 1802)
Drassodes lutescens (C. L. Koch, 1839)
Drassodes pubescens (Thorell, 1856)
Drassodes villosus (Thorell, 1856)
Drassyllus lutetianus (L. Koch, 1866)
Drassyllus praeficus (L. Koch, 1866)
Drassyllus pumilus (C. L. Koch, 1839)
Drassyllus pusillus (C. L. Koch, 1833)
Drassyllus villicus (Thorell, 1875)
Echemus angustifrons (Westring, 1861)
Gnaphosa badia (L. Koch, 1866)
Gnaphosa bicolor (Hahn, 1833)
Gnaphosa dolosa Herman, 1879
Gnaphosa leporina (L. Koch, 1866)
Gnaphosa lucifuga (Walckenaer, 1802)
Gnaphosa lugubris (C. L. Koch, 1839)
Gnaphosa microps Holm, 1939
Gnaphosa modestior Kulczyński, 1897
Gnaphosa moesta Thorell, 1875
Gnaphosa montana (L. Koch, 1866)
Gnaphosa muscorum (L. Koch, 1866)
Gnaphosa opaca Herman, 1879
Gnaphosa rhenana Müller & Schenkel, 1895
Gnaphosa taurica Thorell, 1875
Haplodrassus aenus Thaler, 1984
Haplodrassus bohemicus Miller & Buchar, 1977
Haplodrassus cognatus (Westring, 1861)
Haplodrassus dalmatensis (L. Koch, 1866)
Haplodrassus isaevi Ponomarev & Tsvetkov, 2006
Haplodrassus kulczynskii Lohmander, 1942
Haplodrassus minor (O. Pickard-Cambridge, 1879)
Haplodrassus signifer (C. L. Koch, 1839)
Haplodrassus silvestris (Blackwall, 1833)
Haplodrassus umbratilis (L. Koch, 1866)
Kishidaia conspicua (L. Koch, 1866)
Leptodrassus femineus (Simon, 1873)
Micaria aenea Thorell, 1871
Micaria albovittata (Lucas, 1846)
Micaria coarctata (Lucas, 1846)
Micaria dives (Lucas, 1846)
Micaria formicaria (Sundevall, 1831)
Micaria fulgens (Walckenaer, 1802)
Micaria funerea Simon, 1878
Micaria guttulata (C. L. Koch, 1839)
Micaria pallipes (Lucas, 1846)
Micaria pulicaria (Sundevall, 1831)
Micaria rossica Thorell, 1875
Micaria silesiaca L. Koch, 1875
Micaria sociabilis Kulczyński, 1897
Micaria subopaca Westring, 1861
Nomisia aussereri (L. Koch, 1872)
Nomisia exornata (C. L. Koch, 1839)
Nomisia ripariensis (O. Pickard-Cambridge, 1872)
Phaeocedus braccatus (L. Koch, 1866)
Poecilochroa variana (C. L. Koch, 1839)
Scotophaeus blackwalli (Thorell, 1871)
Scotophaeus quadripunctatus (Linnaeus, 1758)
Scotophaeus scutulatus (L. Koch, 1866)
Setaphis carmeli (O. Pickard-Cambridge, 1872)
Sosticus loricatus (L. Koch, 1866)
Trachyzelotes barbatus (L. Koch, 1866)
Trachyzelotes malkini Platnick & Murphy, 1984
Trachyzelotes pedestris (C. L. Koch, 1837)
Urozelotes rusticus (L. Koch, 1872)
Zelotes aeneus (Simon, 1878)
Zelotes apricorum (L. Koch, 1876)
Zelotes atrocaeruleus (Simon, 1878)
Zelotes aurantiacus Miller, 1967
Zelotes balcanicus Deltshev, 2006 [described from Bulgaria]
Zelotes callidus (Simon, 1878)
Zelotes cingarus (O. Pickard-Cambridge, 1874)
Zelotes clivicola (L. Koch, 1870)
Zelotes electus (C. L. Koch, 1839)
Zelotes erebeus (Thorell, 1871)
Zelotes exiguus (Müller & Schenkel, 1895)
Zelotes gracilis (Canestrini, 1868)
Zelotes harmeron Levy, 2009
Zelotes hermani (Chyzer, 1897)
Zelotes latreillei (Simon, 1878)
Zelotes longipes (L. Koch, 1866)
Zelotes oblongus (C. L. Koch, 1833)
Zelotes petrensis (C. L. Koch, 1839)
Zelotes prope olympi (Kulczyński, 1903)
Zelotes similis (Kulczyński, 1887)
Zelotes strandi (Nosek, 1905)
Zelotes subterraneus (C. L. Koch, 1833)
Zelotes talpinus (L. Koch, 1872)
Zelotes tenuis (L. Koch, 1866)
Zelotes vespertinus (Thorell, 1875)
Zelotes zellensis Grimm, 1982

Family Hahniidae

Antistea elegans (Blackwall, 1841)
Cryphoeca pirini (Drensky, 1921) [described from Bulgaria]
Cryphoeca silvicola (C. L. Koch, 1834)
Hahnia nava (Blackwall, 1841)
Hahnia ononidum Simon, 1875
Hahnia pusilla C. L. Koch, 1841
Hahniharmia picta (Kulczyński, 1897)
Iberina candida (Simon, 1875)
Tuberta maerens (O. Pickard-Cambridge, 1863)

Family Leptonetidae

Cataleptoneta detriticola Deltshev & Li, 2013 [described from Bulgaria]
Protoleptoneta beroni Deltshev, 1977 [described from Bulgaria]
Protoleptoneta bulgarica Deltshev, 1972 [described from Bulgaria]

Family Linyphiidae

Abacoproeces saltuum (L. Koch, 1872)
Acartauchenius scurrilis (O. Pickard-Cambridge, 1872)
Agyneta cauta (O. Pickard-Cambridge, 1902)
Agyneta equestris (L. Koch, 1881)
Agyneta fuscipalpa (C. L. Koch, 1836)
Agyneta gulosa (L. Koch, 1869)
Agyneta ramosa Jackson, 1912
Agyneta rurestris (C. L. Koch, 1836)
Agyneta simplicitarsis (Simon, 1884)
Anguliphantes angulipalpis (Westring, 1851)
Antrohyphantes balcanicus (Drensky, 1931) [described from Bulgaria]
Antrohyphantes rhodopensis (Drensky, 1931) [described from Bulgaria]
Antrohyphantes sophianus (Drensky, 1931) [described from Bulgaria]
Araeoncus anguineus (L. Koch, 1869)
Araeoncus clivifrons Deltshev, 1987 [described from Bulgaria]
Araeoncus crassiceps (Westring, 1861)
Araeoncus humilis (Blackwall, 1841)
Araeoncus tauricus Gnelitsa, 2005
Archaraeoncus prospiciens (Thorell, 1875)
Bathyphantes gracilis (Blackwall, 1841)
Bathyphantes nigrinus (Westring, 1851)
Bolyphantes alticeps (Sundevall, 1833)
Bolyphantes kolosvaryi (Caporiacco, 1936)
Bolyphantes luteolus (Blackwall, 1833)
Caviphantes dobrogicus (Dumitrescu & Miller, 1962)
Centromerita bicolor (Blackwall, 1833)
Centromerus acutidentatus Deltshev, 2002 [described from Bulgaria]
Centromerus brevipalpus (Menge, 1866)
Centromerus bulgarianus (Drensky, 1931) [described from Bulgaria]
Centromerus capucinus (Simon, 1884)
Centromerus cavernarum (L. Koch, 1872)
Centromerus incilium (L. Koch, 1881)
Centromerus lakatnikensis (Drensky, 1931) [described from Bulgaria]
Centromerus milleri Deltshev, 1974 [described from Bulgaria]
Centromerus pabulator (O. Pickard-Cambridge, 1875)
Centromerus prudens (O. Pickard-Cambridge, 1873)
Centromerus semiater (L. Koch, 1879)
Centromerus serratus (O. Pickard-Cambridge, 1875)
Centromerus silvicola (Kulczyński, 1887)
Centromerus sylvaticus (Blackwall, 1841)
Centromerus sylvaticus paucidentatus Deltshev, 1983 [described from Bulgaria]
Centromerus valkanovi Deltshev, 1983 [described from Bulgaria]
Ceratinella brevipes (Westring, 1851)
Ceratinella brevis (Wider, 1834)
Ceratinella major Kulczyński, 1894
Ceratinella scabrosa (O. Pickard-Cambridge, 1871)
Cinetata gradata (Simon, 1881)
Cresmatoneta mutinensis (Canestrini, 1868)
Crosbyarachne silvestris (Georgescu, 1973)
Dicymbium nigrum (Blackwall, 1834)
Dicymbium nigrum brevisetosum Locket, 1962
Dicymbium tibiale (Blackwall, 1836)
Diplocephalus altimontanus Deltshev, 1984 [described from Bulgaria]
Diplocephalus bicephalus (Simon, 1884)
Diplocephalus crassilobus (Simon, 1884)
Diplocephalus cristatus (Blackwall, 1833)
Diplocephalus foraminifer (O. Pickard-Cambridge, 1875)
Diplocephalus graecus (O. Pickard-Cambridge, 1872)
Diplocephalus latifrons (O. Pickard-Cambridge, 1863)
Diplocephalus permixtus (O. Pickard-Cambridge, 1871)
Diplocephalus picinus (Blackwall, 1841)
Diplocephalus protuberans (O. Pickard-Cambridge, 1875)
Diplostyla concolor (Wider, 1834)
Dismodicus bifrons (Blackwall, 1841)
Dismodicus elevatus (C. L. Koch, 1838)
Donacochara speciosa (Thorell, 1875)
Drapetisca socialis (Sundevall, 1833)
Drepanotylus pirinicus Deltshev, 1992 [described from Bulgaria]
Entelecara acuminata (Wider, 1834)
Entelecara flavipes (Blackwall, 1834)
Entelecara media Kulczyński, 1887
Erigone atra Blackwall, 1833
Erigone dentipalpis (Wider, 1834)
Erigone longipalpis pirini Deltshev, 1983 [described from Bulgaria]
Erigonella hiemalis (Blackwall, 1841)
Erigonoplus spinifemuralis Dimitrov, 2003 [described from Bulgaria]
Evansia merens O. Pickard-Cambridge, 1900
Floronia bucculenta (Clerck, 1757)
Frontinellina frutetorum (C. L. Koch, 1834)
Gnathonarium dentatum (Wider, 1834)
Gonatium hilare (Thorell, 1875)
Gonatium nemorivagum (O. Pickard-Cambridge, 1875)
Gonatium orientale Fage, 1931
Gonatium paradoxum (L. Koch, 1869)
Gonatium rubellum (Blackwall, 1841)
Gonatium rubens (Blackwall, 1833)
Gongylidiellum latebricola (O. Pickard-Cambridge, 1871)
Gongylidiellum murcidum Simon, 1884
Gongylidium rufipes (Linnaeus, 1758)
Halorates reprobus (O. Pickard-Cambridge, 1879)
Hilaira excisa (O. Pickard-Cambridge, 1871)
Hylyphantes graminicola (Sundevall, 1830)
Hypomma cornutum (Blackwall, 1833)
Improphantes decolor (Westring, 1861)
Improphantes improbulus (Simon, 1929)
Incestophantes annulatus (Kulczyński, 1882)
Incestophantes crucifer (Menge, 1866)
Ipa keyserlingi (Ausserer, 1867)
Ipa terrenus (L. Koch, 1879)
Kratochviliella bicapitata Miller, 1938
Labulla thoracica (Wider, 1834)
Lasiargus hirsutus (Menge, 1869)
Lepthyphantes centromeroides Kulczyński, 1914
Lepthyphantes leprosus (Ohlert, 1865)
Lepthyphantes notabilis Kulczyński, 1887
Leptorhoptrum robustum (Westring, 1851)
Leptothrix hardyi (Blackwall, 1850)
Lessertia dentichelis (Simon, 1884)
Linyphia hortensis Sundevall, 1830
Linyphia triangularis (Clerck, 1757)
Macrargus carpenteri (O. Pickard-Cambridge, 1894)
Macrargus rufus (Wider, 1834)
Mansuphantes mansuetus (Thorell, 1875)
Mansuphantes prope fragilis (Thorell, 1875)
Mansuphantes rectilamellus (Deltshev, 1988) [described from Bulgaria]
Maso gallicus Simon, 1894
Maso sundevalli (Westring, 1851)
Mecopisthes peusi Wunderlich, 1972
Mecynargus foveatus (Dahl, 1912)
Mecynargus paetulus (O. Pickard-Cambridge, 1875)
Megalepthyphantes collinus (L. Koch, 1872)
Megalepthyphantes nebulosus (Sundevall, 1830)
Metopobactrus orbelicus Deltshev, 1985 [described from Bulgaria]
Micrargus apertus (O. Pickard-Cambridge, 1871)
Micrargus herbigradus (Blackwall, 1854)
Micrargus subaequalis (Westring, 1851)
Microctenonyx subitaneus (O. Pickard-Cambridge, 1875)
Microlinyphia pusilla (Sundevall, 1830)
Microneta viaria (Blackwall, 1841)
Minyriolus pusillus (Wider, 1834)
Moebelia penicillata (Westring, 1851)
Mughiphantes lithoclasicola (Deltshev, 1983) [described from Bulgaria]
Mughiphantes pulcher (Kulczyński, 1881)
Nematogmus sanguinolentus (Walckenaer, 1842)
Neriene clathrata (Sundevall, 1830)
Neriene emphana (Walckenaer, 1842)
Neriene furtiva (O. Pickard-Cambridge, 1871)
Neriene montana (Clerck, 1757)
Neriene peltata (Wider, 1834)
Neriene radiata (Walckenaer, 1842)
Nusoncus nasutus Schenkel 1925
Obscuriphantes obscurus (Blackwall, 1841)
Oedothorax agrestis (Blackwall, 1853)
Oedothorax apicatus (Blackwall, 1850)
Oedothorax fuscus (Blackwall, 1834)
Oedothorax gibbifer (Kulczyński, 1882)
Oedothorax gibbosus (Blackwall, 1841)
Oedothorax retusus (Westring, 1851)
Oreoneta montigena (L. Koch, 1872)
Oreonetides glacialis (L. Koch, 1872)
Ostearius melanopygius (O. Pickard-Cambridge, 1879)
Palliduphantes alutacius (Simon, 1884)
Palliduphantes byzantinus (Fage, 1931)
Palliduphantes insignis (O. Pickard-Cambridge, 1913)
Palliduphantes istrianus (Kulczyński, 1914)
Palliduphantes pallidus (O. Pickard-Cambridge, 1871)
Palliduphantes pillichi (Kulczyński, 1915)
Palliduphantes spelaeorum (Kulczyński, 1914)
Palliduphantes trnovensis (Drensky, 1931) [described from Bulgaria]
Panamomops inconspicuus (Miller & Valesova, 1964)
Panamomops sulcifrons (Wider, 1834)
Pelecopsis elongata (Wider, 1834)
Pelecopsis mengei (Simon, 1884)
Pelecopsis parallela (Wider, 1834)
Pelecopsis radicicola (L. Koch, 1872)
Piniphantes pinicola (Simon, 1884)
Pityohyphantes phrygianus (C. L. Koch, 1836)
Pocadicnemis juncea Locket & Millidge, 1953
Pocadicnemis pumila (Blackwall, 1841)
Poeciloneta variegata (Blackwall, 1841)
Porrhomma convexum (Westring, 1851)
Porrhomma microphthalmum (O. Pickard-Cambridge, 1871)
Porrhomma microps (Roewer, 1931)
Porrhomma pygmaeum (Blackwall, 1834)
Prinerigone vagans (Audouin, 1826)
Saaristoa abnormis (Blackwall, 1841)
Sauron rayi (Simon, 1881)
Scotargus pilosus Simon, 1913
Scotinotylus alpigena (L. Koch, 1869)
Scutpelecopsis krausi (Wunderlich, 1980)
Silometopus bonessi Casemir, 1970
Silometopus reussi (Thorell, 1871)
Sintula retroversus (O. Pickard-Cambridge, 1875)
Sintula spiniger (Balogh, 1935)
Stemonyphantes lineatus (Linnaeus, 1758)
Styloctetor romanus (O. Pickard-Cambridge, 1872)
Syedra gracilis (Menge, 1869)
Tallusia experta (O. Pickard-Cambridge, 1871)
Tallusia vindobonensis (Kulczyński, 1898)
Tapinocyba biscissa (O. Pickard-Cambridge, 1872)
Tapinocyba insecta (L. Koch, 1869)
Tapinocyba mitis (O. Pickard-Cambridge, 1882)
Tapinocyba pallens (O. Pickard-Cambridge, 1872)
Tapinopa longidens (Wider, 1834)
Tenuiphantes alacris (Blackwall, 1853)
Tenuiphantes cristatus (Menge, 1866)
Tenuiphantes drenskyi (van Helsdingen, 1977) [described from Bulgaria]
Tenuiphantes flavipes (Blackwall, 1854)
Tenuiphantes floriana (van Helsdingen, 1977)
Tenuiphantes jacksoni (Schenkel, 1925)
Tenuiphantes jacksonoides (van Helsdingen, 1977)
Tenuiphantes mengei (Kulczyński, 1887)
Tenuiphantes tenebricola (Wider, 1834)
Tenuiphantes tenuis (Blackwall, 1852)
Tenuiphantes zimmermanni (Bertkau, 1890)
Theonina kratochvili Miller & Weiss, 1979
Thyreosthenius biovatus (O. Pickard-Cambridge, 1875)
Thyreosthenius parasiticus (Westring, 1851)
Tiso aestivus (L. Koch, 1872)
Tiso vagans (Blackwall, 1834)
Trichoncoides piscator (Simon, 1884)
Trichoncus affinis Kulczyński, 1894
Trichoncus auritus (L. Koch, 1869)
Trichoncus hackmani Millidge, 1956
Trichoncus saxicola (O. Pickard-Cambridge, 1861)
Trichoncyboides simoni (Lessert, 1904)
Trichopterna cito (O. Pickard-Cambridge, 1872)
Troglohyphantes bureschianus Deltshev, 1975 [described from Bulgaria]
Troglohyphantes drenskii Deltshev, 1973 [described from Bulgaria]
Troxochrus cirrifrons (O. Pickard-Cambridge, 1871)
Troxochrus scabriculus (Westring, 1851)
Typhochrestus digitatus (O. Pickard-Cambridge, 1872)
Walckenaeria acuminata Blackwall, 1833
Walckenaeria alticeps (Denis, 1952)
Walckenaeria antica (Wider, 1834)
Walckenaeria capito (Westring, 1861)
Walckenaeria corniculans (O. Pickard-Cambridge, 1875)
Walckenaeria cucullata (C. L. Koch, 1836)
Walckenaeria dysderoides (Wider, 1834)
Walckenaeria furcillata (Menge, 1869)
Walckenaeria kochi (O. Pickard-Cambridge, 1872)
Walckenaeria mitrata (Menge, 1868)
Walckenaeria monoceros (Wider, 1834)
Walckenaeria nudipalpis (Westring, 1851)
Walckenaeria obtusa Blackwall, 1836
Walckenaeria simplex Chyzer, 1894
Walckenaeria stylifrons (O. Pickard-Cambridge, 1875)
Walckenaeria vigilax (Blackwall, 1853)
Walckenaerianus esyunini Tanasevitch, 2004

Family Liocranidae

Agraecina lineata (Simon, 1878)
Agroeca brunnea (Blackwall, 1833)
Agroeca cuprea Menge, 1873
Agroeca lusatica (L. Koch, 1875)
Agroeca proxima (O. Pickard-Cambridge, 1871)
Apostenus fuscus Westring, 1851
Liocranoeca striata (Kulczyński, 1882)
Liocranum rupicola (Walckenaer, 1830)
Mesiotelus annulipes (Kulczyński, 1897)
Mesiotelus scopensis Drensky, 1935
Mesiotelus tenuissimus (L. Koch, 1866)
Sagana rutilans Thorell, 1875
Scotina celans (Blackwall, 1841)

Family Lycosidae

Alopecosa accentuata (Latreille, 1817)
Alopecosa aculeata (Clerck, 1757)
Alopecosa albofasciata (Brullé, 1832)
Alopecosa cuneata (Clerck, 1757)
Alopecosa cursor (Hahn, 1831)
Alopecosa etrusca Lugetti & Tongiorgi, 1969
Alopecosa fabrilis (Clerck, 1757)
Alopecosa inquilina (Clerck, 1757)
Alopecosa pentheri (Nosek, 1905)
Alopecosa pinetorum (Thorell, 1856)
Alopecosa pulverulenta (Clerck, 1757)
Alopecosa schmidti (Hahn, 1835)
Alopecosa solitaria (Herman, 1879)
Alopecosa striatipes (C. L. Koch, 1839)
Alopecosa sulzeri (Pavesi, 1873)
Alopecosa taeniata (C. L. Koch, 1835)
Alopecosa taeniopus (Kulczyński, 1895)
Alopecosa trabalis (Clerck, 1757)
Arctosa alpigena lamperti Dahl, 1908
Arctosa cinerea (Fabricius, 1777)
Arctosa figurata (Simon, 1876)
Arctosa leopardus (Sundevall, 1833)
Arctosa lutetiana (Simon, 1876)
Arctosa maculata (Hahn, 1822)
Arctosa perita (Latreille, 1799)
Arctosa stigmosa (Thorell, 1875)
Arctosa tbilisiensis Mcheidze, 1946
Arctosa variana C. L. Koch, 1847
Aulonia albimana (Walckenaer, 1805)
Geolycosa vultuosa (C. L. Koch, 1838)
Hogna radiata (Latreille, 1817)
Lycosa praegrandis C. L. Koch, 1836
Lycosa singoriensis (Laxmann, 1770)
Pardosa agrestis (Westring, 1861)
Pardosa agricola (Thorell, 1856)
Pardosa alacris (C. L. Koch, 1833)
Pardosa albatula (Roewer, 1951)
Pardosa amentata (Clerck, 1757)
Pardosa atomaria (C. L. Koch, 1847)
Pardosa bifasciata (C. L. Koch, 1834)
Pardosa blanda (C. L. Koch, 1833)
Pardosa consimilis Nosek, 1905
Pardosa cribrata Simon, 1876
Pardosa drenskii Buchar, 1968
Pardosa ferruginea (L. Koch, 1870)
Pardosa hortensis (Thorell, 1872)
Pardosa incerta Nosek, 1905
Pardosa italica Tongiorgi, 1966
Pardosa luctinosa Simon, 1876
Pardosa lugubris (Walckenaer, 1802)
Pardosa mixta (Kulczyński, 1887)
Pardosa monticola (Clerck, 1757)
Pardosa morosa (L. Koch, 1870)
Pardosa nebulosa (Thorell, 1872)
Pardosa nigra (C. L. Koch, 1834)
Pardosa nigriceps (Thorell, 1856)
Pardosa paludicola (Clerck, 1757)
Pardosa palustris (Linnaeus, 1758)
Pardosa pontica (Thorell, 1875)
Pardosa prativaga (L. Koch, 1870)
Pardosa proxima (C. L. Koch, 1847)
Pardosa pseudostrigillata Tongiorgi, 1966
Pardosa pullata (Clerck, 1757)
Pardosa riparia (C. L. Koch, 1833)
Pardosa roscai (Roewer, 1951)
Pardosa tasevi Buchar, 1968
Pardosa tatarica (Thorell, 1875)
Pardosa vittata (Keyserling, 1863)
Pirata piraticus (Clerck, 1757)
Pirata piscatorius (Clerck, 1757)
Pirata tenuitarsis Simon, 1876
Piratula hygrophila (Thorell, 1872)
Piratula insularis (Emerton, 1885)
Piratula knorri (Scopoli, 1763)
Piratula latitans (Blackwall, 1841)
Trabea paradoxa Simon, 1876
Trochosa hispanica Simon, 1870
Trochosa robusta (Simon, 1876)
Trochosa ruricola (De Geer, 1778)
Trochosa terricola Thorell, 1856
Xerolycosa miniata (C. L. Koch, 1834)
Xerolycosa nemoralis (Westring, 1861)

Family Mimetidae

Ero aphana (Walckenaer, 1802)
Ero cambridgei Kulczyński, 1911
Ero furcata (Villers, 1789)
Ero tuberculata (De Geer, 1778)
Mimetus laevigatus (Keyserling, 1863)

Family Miturgidae

Zora armillata Simon, 1878
Zora manicata Simon, 1878
Zora nemoralis (Blackwall, 1861)
Zora parallela Simon, 1878
Zora pardalis Simon, 1878
Zora silvestris Kulczyński, 1897
Zora spinimana (Sundevall, 1833)

Family Nemesiidae

Brachythele denieri (Simon, 1916)
Brachythele langourovi Lazarov, 2005 [described from Bulgaria]
Nemesia caementaria (Latreille, 1799)
Nemesia pannonica coheni Fuhn & Polenec, 1967

Family Nesticidae

Nesticus beroni Deltshev, 1977 [described from Bulgaria]
Nesticus cellulanus (Clerck, 1757)
Nesticus eremita Simon, 1879

Family Oecobiidae

Uroctea durandi (Latreille, 1809)

Family Oonopidae

Oonops pulcher Templeton, 1835
Orchestina pavesii (Simon, 1873)
Silhouettella loricatula (Roewer, 1942)
Tapinesthis inermis (Simon, 1882)

Family Oxyopidae

Oxyopes heterophthalmus (Latreille, 1804)
Oxyopes lineatus Latreille, 1806
Oxyopes ramosus (Martini & Goeze, 1778)

Family Philodromidae

Philodromus albidus Kulczyński, 1911
Philodromus aureolus (Clerck, 1757)
Philodromus buxi Simon, 1884
Philodromus cespitum (Walckenaer, 1802)
Philodromus collinus C. L. Koch, 1835
Philodromus dispar Walckenaer, 1826
Philodromus emarginatus (Schrank, 1803)
Philodromus fallax Sundevall, 1833
Philodromus fuscomarginatus (De Geer, 1778)
Philodromus histrio (Latreille, 1819)
Philodromus longipalpis Simon, 1870
Philodromus margaritatus (Clerck, 1757)
Philodromus marmoratus Kulczyński, 1891
Philodromus monitae Muster, 2010
Philodromus pinetorum Muster, 2009
Philodromus poecilus (Thorell, 1872)
Philodromus praedatus O. Pickard-Cambridge, 1871
Philodromus rufus Walckenaer, 1826
Philodromus vagulus Simon, 1875
Pulchellodromus glaucinus (Simon, 1870)
Thanatus arenarius L. Koch, 1872
Thanatus atratus Simon, 1875
Thanatus flavidus Simon, 1875
Thanatus formicinus (Clerck, 1757)
Thanatus imbecillus L. Koch, 1878
Thanatus lineatipes Simon, 1870
Thanatus meronensis Levy, 1977
Thanatus oblongiusculus (Lucas, 1846)
Thanatus pictus L. Koch, 1881
Thanatus sabulosus (Menge, 1875)
Thanatus striatus C. L. Koch, 1845
Thanatus vulgaris Simon, 1870
Thanatus vulgaris creticus Kulczyński, 1903
Tibellus macellus Simon, 1875
Tibellus maritimus (Menge, 1875)
Tibellus oblongus (Walckenaer, 1802)

Family Pholcidae

Holocnemus pluchei (Scopoli, 1763)
Hoplopholcus forskali (Thorell, 1871)
Hoplopholcus labyrinthi (Kulczyński, 1903)
Pholcus opilionoides (Schrank, 1781)
Pholcus phalangioides (Fuesslin, 1775)
Pholcus ponticus Thorell, 1875
Psilochorus simoni (Berland, 1911)
Spermophora senoculata (Dugès, 1836)

Family Phrurolithidae

Phrurolithus festivus (C. L. Koch, 1835)
Phrurolithus pullatus Kulczyński, 1897
Phrurolithus szilyi Herman, 1879

Family Pisauridae

Dolomedes fimbriatus (Clerck, 1757)
Dolomedes plantarius (Clerck, 1757)
Pisaura mirabilis (Clerck, 1757)

Family Salticidae

Aelurillus v-insignitus (Clerck, 1757)
Asianellus festivus (C. L. Koch, 1834)
Ballus chalybeius (Walckenaer, 1802)
Carrhotus xanthogramma (Latreille, 1819)
Chalcoscirtus infimus (Simon, 1868)
Chalcoscirtus nigritus (Thorell, 1875)
Chalcoscirtus pseudoinfimus Ovtsharenko, 1978
Cyrba algerina (Lucas, 1846)
Dendryphantes hastatus (Clerck, 1757)
Dendryphantes rudis (Sundevall, 1833)
Euophrys frontalis (Walckenaer, 1802)
Euophrys herbigrada (Simon, 1871)
Euophrys rufibarbis (Simon, 1868)
Evarcha arcuata (Clerck, 1757)
Evarcha falcata (Clerck, 1757)
Evarcha jucunda (Lucas, 1846)
Evarcha laetabunda (C. L. Koch, 1846)
Heliophanus aeneus (Hahn, 1832)
Heliophanus auratus C. L. Koch, 1835
Heliophanus cupreus (Walckenaer, 1802)
Heliophanus dampfi Schenkel, 1923
Heliophanus dubius C. L. Koch, 1835
Heliophanus flavipes (Hahn, 1832)
Heliophanus kochii Simon, 1868
Heliophanus lineiventris Simon, 1868
Heliophanus melinus L. Koch, 1867
Heliophanus patagiatus Thorell, 1875
Heliophanus rufithorax Simon, 1868
Heliophanus simplex Simon, 1868
Heliophanus tribulosus Simon, 1868
Leptorchestes berolinensis (C. L. Koch, 1846)
Leptorchestes mutilloides (Lucas, 1846)
Macaroeris flavicomis (Simon, 1884)
Macaroeris nidicolens (Walckenaer, 1802)
Marpissa muscosa (Clerck, 1757)
Marpissa nivoyi (Lucas, 1846)
Marpissa pomatia (Walckenaer, 1802)
Marpissa radiata (Grube, 1859)
Mendoza canestrinii (Ninni, 1868)
Menemerus falsificus Simon, 1868
Menemerus semilimbatus (Hahn, 1829)
Menemerus taeniatus (L. Koch, 1867)
Mogrus neglectus (Simon, 1868)
Myrmarachne formicaria (De Geer, 1778)
Neon levis (Simon, 1871)
Neon pictus Kulczyński, 1891
Neon rayi (Simon, 1875)
Neon reticulatus (Blackwall, 1853)
Pellenes arciger (Walckenaer, 1837)
Pellenes diagonalis (Simon, 1868)
Pellenes geniculatus (Simon, 1868)
Pellenes nigrociliatus (Simon, 1875)
Pellenes seriatus (Thorell, 1875)
Pellenes tripunctatus (Walckenaer, 1802)
Philaeus chrysops (Poda, 1761)
Phintella castriesiana (Grube, 1861)
Phlegra bresnieri (Lucas, 1846)
Phlegra fasciata (Hahn, 1826)
Plexippoides gestroi (Dalmas, 1920)
Pseudeuophrys erratica (Walckenaer, 1826)
Pseudeuophrys lanigera (Simon, 1871)
Pseudeuophrys obsoleta (Simon, 1868)
Pseudicius badius (Simon, 1868)
Pseudicius encarpatus (Walckenaer, 1802)
Pseudicius kulczynskii Nosek, 1905
Pseudicius picaceus (Simon, 1868)
Saitis barbipes (Simon, 1868)
Saitis graeca Kulczyński, 1905
Saitis taurica Kulczyński, 1905
Salticus cingulatus (Panzer, 1797)
Salticus confusus Lucas, 1846
Salticus mutabilis Lucas, 1846
Salticus scenicus (Clerck, 1757)
Salticus zebraneus (C. L. Koch, 1837)
Sibianor aurocinctus (Ohlert, 1865)
Sitticus atricapillus (Simon, 1882)
Sitticus damini (Chyzer, 1891)
Sitticus distinguendus (Simon, 1868)
Sitticus dzieduszyckii (L. Koch, 1870)
Sitticus floricola (C. L. Koch, 1837)
Sitticus inexpectus Logunov & Kronestedt, 1997
Sitticus penicillatus (Simon, 1875)
Sitticus pubescens (Fabricius, 1775)
Sitticus rupicola (C. L. Koch, 1837)
Sitticus saltator (O. Pickard-Cambridge, 1868)
Sitticus saxicola (C. L. Koch, 1846)
Sitticus terebratus (Clerck, 1757)
Synageles dalmaticus (Keyserling, 1863)
Synageles hilarulus (C. L. Koch, 1846)
Synageles venator (Lucas, 1836)
Talavera aequipes (O. Pickard-Cambridge, 1871)
Talavera monticola (Kulczyński, 1884)
Talavera petrensis (C. L. Koch, 1837)
Yllenus arenarius Menge, 1868
Yllenus horvathi Chyzer, 1891

Family Scytodidae

Scytodes thoracica (Latreille, 1802)

Family Segestriidae

Segestria bavarica C. L. Koch, 1843
Segestria florentina (Rossi, 1790)
Segestria senoculata (Linnaeus, 1758)

Family Sparassidae

Micrommata ligurina (C. L. Koch, 1845)
Micrommata virescens (Clerck, 1757)
Olios argelasius (Walckenaer, 1805)

Family Tetragnathidae

Meta bourneti Simon, 1922
Meta menardi (Latreille, 1804)
Metellina mengei (Blackwall, 1870)
Metellina merianae (Scopoli, 1763)
Metellina segmentata (Clerck, 1757)
Pachygnatha clercki Sundevall, 1823
Pachygnatha clerckoides Wunderlich, 1985
Pachygnatha degeeri Sundevall, 1830
Pachygnatha listeri Sundevall, 1830
Tetragnatha dearmata Thorell, 1873
Tetragnatha extensa (Linnaeus, 1758)
Tetragnatha montana Simon, 1874
Tetragnatha nigrita Lendl, 1886
Tetragnatha nitens (Audouin, 1826)
Tetragnatha obtusa C. L. Koch, 1837
Tetragnatha pinicola L. Koch, 1870
Tetragnatha striata L. Koch, 1862

Family Theridiidae

Anelosimus vittatus (C. L. Koch, 1836)
Asagena meridionalis Kulczyński, 1894
Asagena phalerata (Panzer, 1801)
Crustulina guttata (Wider, 1834)
Crustulina scabripes Simon, 1881
Crustulina sticta (O. Pickard-Cambridge, 1861)
Dipoena braccata (C. L. Koch, 1841)
Dipoena coracina (C. L. Koch, 1837)
Dipoena croatica (Chyzer, 1894)
Dipoena erythropus Simon (1881)
Dipoena melanogaster (C. L. Koch, 1837)
Dipoena nigroreticulata (Simon, 1879)
Dipoena torva (Thorell, 1875)
Enoplognatha afrodite Hippa & Oksala, 1983
Enoplognatha latimana Hippa & Oksala, 1982
Enoplognatha mandibularis (Lucas, 1846)
Enoplognatha mordax (Thorell, 1875)
Enoplognatha oelandica (Thorell, 1875)
Enoplognatha ovata (Clerck, 1757)
Enoplognatha penelope Hippa & Oksala, 1982
Enoplognatha quadripunctata Simon, 1884
Enoplognatha thoracica (Hahn, 1833)
Episinus angulatus (Blackwall, 1836)
Episinus maculipes Cavanna, 1876
Episinus truncatus Latreille, 1809
Euryopis episinoides (Walckenaer, 1847)
Euryopis flavomaculata (C. L. Koch, 1836)
Euryopis laeta (Westring, 1861)
Euryopis quinqueguttata Thorell, 1875
Euryopis saukea Levi, 1951
Euryopis sexalbomaculata (Lucas, 1846)
Heterotheridion nigrovariegatum (Simon, 1873)
Kochiura aulica (C. L. Koch, 1838)
Lasaeola convexa (Blackwall, 1870)
Lasaeola prona (Menge, 1868)
Lasaeola tristis (Hahn, 1833)
Latrodectus tredecimguttatus (Rossi, 1790)
Neottiura bimaculata (Linnaeus, 1767)
Neottiura herbigrada (Simon, 1873)
Neottiura suaveolens (Simon, 1879)
Neottiura uncinata (Lucas, 1846)
Ohlertidion ohlerti (Thorell, 1870)
Paidiscura pallens (Blackwall, 1834)
Parasteatoda lunata (Clerck, 1757)
Parasteatoda simulans (Thorell, 1875)
Parasteatoda tabulata (Levi, 1980)
Parasteatoda tepidariorum (C. L. Koch, 1841)
Pholcomma gibbum (Westring, 1851)
Phycosoma inornatum (O. Pickard-Cambridge, 1861)
Phylloneta impressa (L. Koch, 1881)
Phylloneta sisyphia (Clerck, 1757)
Platnickina tincta (Walckenaer, 1802)
Robertus arundineti (O. Pickard-Cambridge, 1871)
Robertus frivaldszkyi (Chyzer, 1894)
Robertus lividus (Blackwall, 1836)
Robertus mediterraneus Eskov, 1987
Robertus neglectus (O. Pickard-Cambridge, 1871)
Robertus scoticus Jackson, 1914
Rugathodes bellicosus (Simon, 1873)
Sardinidion blackwalli O. Pickard-Cambridge, 1871
Simitidion simile (C. L. Koch, 1836)
Steatoda albomaculata (De Geer, 1778)
Steatoda bipunctata (Linnaeus, 1758)
Steatoda castanea (Clerck, 1757)
Steatoda grossa (C. L. Koch, 1838)
Steatoda paykulliana (Walckenaer, 1805)
Steatoda triangulosa (Walckenaer, 1802)
Theridion adrianopoli Drensky, 1915 [described from Bulgaria]
Theridion betteni Wiehle, 1960
Theridion boesenbergi Strand, 1904
Theridion cinereum Thorell, 1875
Theridion hemerobium Simon, 1914
Theridion melanurum Hahn, 1831
Theridion mystaceum L. Koch, 1870
Theridion petraeum L. Koch, 1872
Theridion pictum (Walckenaer, 1802)
Theridion pinastri L. Koch, 1872
Theridion varians Hahn, 1833

Family Theridiosomatidae

Theridiosoma gemmosum (L. Koch, 1877)

Family Thomisidae

Coriarachne depressa (C. L. Koch, 1837)
Cozyptila blackwalli (Simon, 1875)
Cozyptila thaleri Marusik & Kovblyuk, 2005
Diaea dorsata (Fabricius, 1777)
Diaea livens Simon, 1876
Ebrechtella tricuspidata (Fabricius, 1775)
Heriaeus graminicola (Doleschall, 1852)
Heriaeus hirtus (Latreille, 1819)
Heriaeus mellotteei Simon, 1886
Heriaeus setiger (O. Pickard-Cambridge, 1872)
Heriaeus simoni Kulczyński, 1903
Misumena vatia (Clerck, 1757)
Monaeses paradoxus (Lucas, 1846)
Ozyptila atomaria (Panzer, 1801)
Ozyptila brevipes (Hahn, 1826)
Ozyptila claveata (Walckenaer, 1837)
Ozyptila confluens (C. L. Koch, 1845)
Ozyptila conostyla Hippa, Koponen & Oksala, 1986
Ozyptila gertschi Kurata, 1944
Ozyptila lugubris (Kroneberg, 1875)
Ozyptila praticola (C. L. Koch, 1837)
Ozyptila prope umbraculorum Simon, 1932
Ozyptila pullata (Thorell, 1875)
Ozyptila rauda Simon, 1875
Ozyptila sanctuaria (O. Pickard-Cambridge, 1871)
Ozyptila scabricula (Westring, 1851)
Ozyptila simplex (O. Pickard-Cambridge, 1862)
Ozyptila trux (Blackwall, 1846)
Pistius truncatus (Pallas, 1772)
Runcinia grammica (C. L. Koch, 1837)
Synema globosum (Fabricius, 1775)
Synema plorator (O. Pickard-Cambridge, 1872)
Thomisus onustus Walckenaer, 1805
Tmarus piger (Walckenaer, 1802)
Tmarus piochardi (Simon, 1866)
Tmarus stellio Simon, 1875
Xysticus abditus Logunov, 2006 [described from Bulgaria]
Xysticus acerbus Thorell, 1872
Xysticus audax (Schrank, 1803)
Xysticus bifasciatus C. L. Koch, 1837
Xysticus bonneti Denis, 1938
Xysticus caperatus Simon, 1875
Xysticus cor Canestrini, 1873
Xysticus cristatus (Clerck, 1757)
Xysticus desidiosus Simon, 1875
Xysticus erraticus (Blackwall, 1834)
Xysticus gallicus Simon, 1875
Xysticus graecus C. L. Koch, 1837
Xysticus kempeleni Thorell, 1872
Xysticus kochi Thorell, 1872
Xysticus laetus Thorell, 1875
Xysticus lanio C. L. Koch, 1835
Xysticus lineatus (Westring, 1851)
Xysticus luctator L. Koch, 1870
Xysticus luctuosus (Blackwall, 1836)
Xysticus macedonicus Silhavy, 1944
Xysticus marmoratus Thorell, 1875
Xysticus ninnii Thorell, 1872
Xysticus nubilus Simon, 1875
Xysticus robustus (Hahn, 1832)
Xysticus sabulosus (Hahn, 1832)
Xysticus striatipes L. Koch, 1870
Xysticus tenebrosus Silhavy, 1944
Xysticus ulmi (Hahn, 1831)

Family Titanoecidae

Nurscia albomaculata (Lucas, 1846)
Nurscia albosignata Simon, 1874
Titanoeca incerta (Nosek, 1905)
Titanoeca quadriguttata (Hahn, 1833)
Titanoeca schineri L. Koch, 1872
Titanoeca tristis L. Koch, 1872
Titanoeca veteranica Herman, 1879

Family Trachelidae

Cetonana laticeps (Canestrini, 1868)
Metatrachelas rayi (Simon, 1878)
Paratrachelas maculatus (Thorell, 1875)

Family Uloboridae

Hyptiotes flavidus (Blackwall, 1862)
Hyptiotes paradoxus (C. L. Koch, 1834)
Uloborus plumipes Lucas, 1846
Uloborus walckenaerius Latreille, 1806

Family Zodariidae

Zodarion aculeatum Chyzer, 1897
Zodarion blagoevi Bosmans, 2009 [described from Bulgaria]
Zodarion epirense Brignoli, 1984
Zodarion frenatum Simon, 1884
Zodarion graecum (C. L. Koch, 1843)
Zodarion italicum (Canestrini, 1868)
Zodarion morosum Denis, 1935
Zodarion ohridense Wunderlich, 1973
Zodarion pirini Drensky, 1921 [described from Bulgaria]
Zodarion thoni Nosek, 1905
Zodarion turcicum Wunderlich, 1980

Family Zoropsidae

Zoropsis lutea (Thorell, 1875)
Zoropsis spinimana (Dufour, 1820)

Acknowledgements


We are much obliged to all colleagues, who helped us providing references and information on the distribution of the spiders in Bulgaria. A special thank goes to Stanislav Abadjiev (lepidopterologist, NMNHS) who arranged the new version of the site. We are very grateful also to Peter van Helsdingen, Wolfgang Nentwig and collaborators and to Norman Platnick for making available to all arachnologists the websites ‘Fauna Europaea’, ‘Spiders of Europe’ and ‘The World Spider Catalog’.

References


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Deltshev C. 1995b. Spiders (Araneae) from the high altitude zone of Rila Mountain (Bulgaria). — Ber. naturw.-med. Ver. Innsbruck, 82: 217-225.
Deltshev C. 1996. The origin, formation and zoogeography of endemic spiders of Bulgaria (Araneae). — Revue suisse Zool., vol. hors serie 1: 141-151.
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Deltshev C. 1997b. Spiders (Araneae) from the coastal habitats of Shabla-Ezerets Lake, Bulgaria. — Acta zool. bulg., 49: 58-63.
Deltshev C. 1998. Spiders from the high altitude zone of Central Stara Planina Mountain (Bulgaria) (Araneae). — Ber. naturw.-med. Ver. Innsbruck, 85: 213-221.
Deltshev C. 1999. Faunistic and zoogeographical review of the Spiders (Araneae) of the Balkan peninsula. — The Journal of Arachnology. Kansas, 27: 255-261.
Deltshev C. 2000. The endemic spiders (Araneae) of the Balkan Peninsula. — In: Gajdos, P., Pekar, S. (Eds), Proceedings of the 18th European Colloquium of Arachnology, Stara Lesna, 1999. Ekolуgia (Bratislava), 19(3): 59-65.
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Deltshev C. 2003b. A Zoogeographical review of Spiders (Araneae) in Balkan Peninsula. — In: Proceedings of Koper, Klower, Amsterdam.
Deltshev C. 2003c. Faunistic Diversity and Conservation Significanse of Spiders (Araneae). — In: Dimitar Peev (Ed.), Rapid Ecological Assessment of Rila Monastery Nature Park, USAID, 108-116.
Deltshev C. 2004. Spiders (Araneae) from Sandanski-Petrich Valley (SW Bulgaria). — Mitt. Mus. Nat. kd. Berl., Zool. Reihe, 80 (1): 71-76.
Deltshev C. 2005. A new Hypomma species from Stara Planina Mountains, Bulgaria (Araneae, Linyphiidae). — Revue suisse de Zoologie 112 (1): 115-119.
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Deltshev C. 2008. Two new spider species, Malthonica bozhkovi sp. nov. and Tegenaria paragamiani sp. nov. from Rhodopy Mountains (Bulgaria and Greece) (Araneae: Agelenidae). — Zootaxa, 1872: 37-44.
Deltshev C. 2011a. Faunistic Diversity of Spiders (Araneae) in Castanian forests of Belasitsa Mt. — Report. State and prospects of the Castanea sativa population in Belasitsa mountain: climate change adaptation; maintenance of biodiversity and sustainable ecosystem management: 1-12.
Deltshev C. 2011b. Harpactea lazarovi sp.n. and H. tenuiemboli sp.tr., two new spider species from Balkan Peninsula (Araneae, Dysderidae). — Zoosystematics and Evolution, 87 (2): 221-226.
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The Spiders (Araneae) of Bulgaria

This check list should be cited as follows:
Blagoev, G., Deltshev, C. & Lazarov, S. 2002—2017. The Spiders (Araneae) of Bulgaria. — National Museum of Natural History, Bulgarian Academy of Sciences, online at http://www.nmnhs.com/spiders-bulgaria/