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Staff publications

Nikolai Spassov [publications]

Spassov, N., Iliev, N., (in press). Notes on the animal bone remains from the early excavations (1956—1959) of the late Chalcolithic site near Hotnitsa. — In: St. Chohadjiev (ed.) Hotnitsa — The Prehistoric settlement, Vol. I. Tirnovo.

Spassov, N., Iliev, N., (in press). Analyses of the bones from domestic and wild mammals from the Bronze Age settlement near Baley. — In: St. Alexandrov (ed.). Baley, the settlement from the Late Bronze age. Sofia.

Spassov, N., Georgiev, K., Ivanova, N., Ivanov, V., (in press). Study on the status of the marbled polecat (Vormela peregusna peregusna Guld.) in Southern Dobrudja (Bulgaria). — Bull. of the Mustelidae, Viverridae & Procyonidae group IUCN/SSC.

Spassov, N., Georgiev, K., Ivanov, V., (in press). The Population Status and Dynamics of the Brown Bear (Ursus arctos L.) on the Cross-Border Region between Bulgaria and Greece. — Bull. of the Belgrade Mus. of Nat. Hist.

Спасов, Н., Илиев Н., Боев, З., (in press). Животинските останки от ранно-неолитното селище край с. Капитан-Димитриево ( Пазарджишка област). — В: В. Николов и др. Селищна могила Капитан Димитриево. Разкопки 2001—2005. Под ред. на В. Николов. София, 2011.

Yankova, I., Marinov, M., Neov, B., Petrova, M., Spassov, N., Hristov, P., Radoslavov, G., 2019. Evidence for Early European Neolithic Dog Dispersal: New Data on Southeastern European Subfossil Dogs from the Prehistoric and Antiquity Ages. — Genes, 10 (10): 757. DOI: 10.3390/genes10100757

Spassov, N., Geraads, D., Hristova, L., Markov, G. N., 2019. The Late Miocene (Turolian) mammal fauna from Gorna Sushitsa, Southwestern Bulgaria. — Neues Jahrbuch für Geologie und Paläontologie, 291 (3): 317—350.

Spassov, N., Acosta-Pankov, I., 2019. Dispersal history of the golden jackal (Canis aureus moreoticus Geoffroy, 1835) in Europe and possible causes of its recent population explosion. — Biodiversity Data Journal, 7: e34825. DOI: 10.3897/BDJ.7.e34825

Nishita, Y., Spassov, N., Peeva, S., Raichev, E. G., Kaneko, Y., Masuda, R., 2019. Genetic diversity of MHC class II DRB alleles in the marbled polecat, Vormela peregusna, in Bulgaria. — Ethology Ecology & Evolution, 31 (1): 59—72. DOI: 10.1080/03949370.2018.1486887

Ersmark, E., Baryshnikov, G., Higham, T., Argant, A., Castaños, P., Döppes, D., Gasparik, M., Germonpré, M., Lidén, K., Lipecki, G., Marciszak, A., Miller, R., Moreno‐García, M., Pacher, M., Robu, M., Rodriguez-Manuel Rojo Guerra, R., Sabol, M., Spassov, N., Storå, J., Valdiosera, C., Villaluenga, A., Stewart, J. R., Dalén, L., 2019. Genetic turnovers and northern survival during the last glacial maximum in European brown bears. — Ecology and Evolution, 9 (10): 5891—5905. DOI: 10.1002/ece3.5172

Böhme, M., Spassov, N., Fuss, J., Tröscher, A., Deane, A. S., Prieto, J., Kirscher, U., Lechner, T., Begun, D. R., 2019. A new Miocene ape and locomotion in the ancestor of great apes and humans. — Nature. DOI: 10.1038/s41586-019-1731-0

Spassov, N., Iliev, N., Hristova, L., Ivanov, V., 2018. Typology of Thracian horses according to osteological analysis of skeletal remains and depictions from the antiquity. — Historia naturalis bulgarica, 25: 15—24.

Spassov, N., Hristova, L., Iliev, N., 2018. The domesticated horses from the submerged prehistoric village of Urdoviza (Kiten) on the Bulgarian Black Sea coast — among the oldest known. — Historia naturalis bulgarica, 25: 11—14.

Spassov, N., Hristova, L., 2018. Early Ursus etruscus Cuvier from the Middle Villafranchian carst cavern locality of Varshets (Bulgaria). — 24th International Cave Bear Symposium 2018, Chepelare, 27—30.09.2018.

Spassov, N., Geraads, D., Hristova, L., Markov, G. N., Garevska, B., Garevski, R., 2018. The late Miocene mammal faunas of the Republic of Macedonia (FYROM). — Palaeontographica A., 311 (1—6): 1—85.

Mennecart, B., Geraads, D., Spassov, N., Zagorchev, I., 2018. Discovery of the oldest European ruminant in the late Eocene of Bulgaria: Did tectonics influence the diachronic development of the Grande Coupure? — Palaeogeography, Palaeoclimatology, Palaeoecology, 498: 1—8.

Gurova, M., Ivanova, S., Marinova, E., V., Spassov, N., Hristova, L., Verheyden, S., Burlet, C., 2018. Excavations at Mishin Kamik Cave: fifth season. — Archaeological Discoveries and Excavations in 2017, National Archaeological Institute with Museum at the Bulgarian Academy of Sciences,8—11.

Fuss, J., Spassov, N., Böhme, M., Begun, D. R., 2018. Response to Benoit and Thackeray ‘A cladistic analysis of Graecopithecus’. — South African Journal of Science, 114 (5/6): Art. #a0267, 2 pp. DOI: 10.17159/sajs.2018/a0267

Dzhebir, G., Yordanov, G., Yankova, I., Sirakova, D., Petrova, M., Neov, B., Hristov, P., Radoslavov, G., Hristova, L., Spassov, N., 2018. Comparative genetic analysis of subfossil wild horses (from the Neolithic Age and Early Bronze Age) and present-day domestic horses from Bulgaria. — Historia naturalis bulgarica, 25: 3—10.

Dzhebir, G., Yordanov, G., Yankova, I., Sirakova, D., Petrova, M., Neov, B., Hristov, P., Radoslavov, G., Hristova, L., Spassov, N., 2018. Comparative genetic analysis of modern and ancient horses from the Neolithic age in Bulgaria. — Historia naturalis bugarica, 28: 3—10.

Böhme, M., Van Baak, C. G., Prieto, J., Winklhofer, M., Spassov, N., 2018. Late Miocene stratigraphy, palaeoclimate and evolution of the Sandanski Basin (Bulgaria) and the chronology of the Pikermian faunal changes. — Global and planetary change, 170: 1—19.

Acosta-Pankov, I., Spassov, N., Banea, O. C., 2018. Seasonal Differences in Howling Response of Golden Jackals Canis aureus L. 1758 (Mammalia: Canidae) in Eastern Bulgaria. — Acta zoologica bulgarica, 70 (4): 539—545.

Spassov, N., Hristova, L., Ivanova, S., Georgiev, I., 2017. First record of the small cave bear in Bulgaria and the taxonomic status of bears of the Ursus savini Andrews — Ursus rossicus Borissiak group. — Fossil imprint, 73 (3—4): 275—291.

Spassov, N., Hristova, L., Ivanova, S., Georgiev, I., 2017. First Record of the ‘Small Cave Bear’ in Bulgaria and taxonomic status of the bears of the Ursus savini Andrews — Ursus rossicus Borissiak group. — Fossil Imprint, 73 (3—4): 275—291.

Spassov, N., Böhme, M., Geraads, D., Kötter, S., Van Baak, C., 2017. Pikermian mammal event, post-Pikermian mammal turnover and appearance of Graecopithecus. — 5th Congress of the RCMNS. September 03—06.2017. Book of Abstracts, p. 21.

Ottoni, C., Van Neer, W., De Cupere, B., Daligault, J., Guimaraes, S., Peters, J., Spassov, N., Prendergast, M. E., Boivin, N., Morales-Muñiz, A., Bălăşescu, A., Becker, C., Benecke, N., Boroneant, A., Buitenhuis, H., Chahoud, J., Crowther, A., Llorente, L., Manaseryan, N., Monchot, H., Onar, V., Osypińska, M., Putelat, O., Quintana Morales, E. M., Studer, J., Wierer, U., Decorte, R., Grange, T., Geigl, E.-M., 2017. The palaeogenetics of cat dispersal in the ancient world. — Nature Ecology & Evolution, 1 (7): 1—7.

Mizumachi, K., Nishita, Y., Spassov, N., Raichev, E. G., Peeva, S., Kaneko, Y., Masuda, R., 2017. Molecular phylogenetic status of the Bulgarian marbled polecat (Vormela peregusna, Mustelidae, Carnivora), revealed by Y chromosomal genes and mitochondrial DNA sequences. — Biochemical Systematics and Ecology, 70: 99—107. DOI: 10.1016/j.bse.2016.10.025

Milchev, B., Spassov, N., 2017. First evidence for carrion–feeding of Eurasian Eagle-owl (Bubo bubo) in Bulgaria. — Ornis Hungarica, 25 (1): 58—69.

Li, Y., Spassov, N., 2017. A new species of Paramachaerodus (Mammalia, Carnivora, Felidae) from the late Miocene of China and Bulgaria, and revision of Promegantereon Kretzoi, 1938 and Paramachaerodus Pilgrim, 1913. — Paläontologische Zeitschrift, 3 (91): 409—426. DOI: 10.1007/s12542-017-0371-7

Gurova, M., Ivanova, S., Spassov, N., Hristova, L., Popov, V., Marinova, E., Böhme, M., 2017. Excavations at Mishin Kamik Cave: fourth season. 2016 season. — Archaeological Discoveries and Excavations in 2016, National Archaeological Institute with Museum at the Bulgarian Academy of Sciences, 48—50.

Gurov, T., Atanassov, E., Karaivanova, A., Serbezov, R., Spassov, N., 2017. Statistical Estimation of Brown Bears (Ursus arctos L.) Population in the Rhodope mountains. — Procedia Computer Science 108C (2017): 2028—2037.

Fuss, J., Spassov, N., Begun, D., Böhme, M., 2017. Potential hominin affinities of Graecopithecus from the Late Miocene of Europe. — PlosOne, 12: e0177127.

Böhme, M., Spassov, N., Ebner, M., Geraads, D., Hristova, L., Kirscher, U., Kötter, S., Linnemann, U., Prieto, J., Roussiakis, S., Theodorou, G., Uhlig, G., Winklhofer, M., 2017. Messinian Age and savanna environment of the possible hominin Graecopithecus from SE Europe. — PlosOne, 12 (5): e0177347. DOI: 10.1371/journal.pone.0177347

Böhme, M., Spassov, N., Ebner, M., Geraads, D., Hristova, L., Kirscher, U., Kötter, S., Linnemann, U., Prieto, J., Roussiakis, S., Theodorou, G., Uhlig, G., Winklhofer, M., 2017. Messinian Age and savanna environment of the possible hominin Graecopithecus from SE Europe. — PLoS One, 12 (5): e0177347. DOI: 10.1371/journal.pone.0177347

Böhme, M., Spassov, N., Ebner, M., Geraads, D., Hristova, L., Kirscher, U., Kötter, S., Linnemann, U., Prieto, J., Roussiakis, S., Theodorou, G., Uhlig, G., Winklhofer, M., 2017. Environment and stratigraphy of the classical Pikermi sites and post-Pikermian localities and the first, Late Miocene, Sahara desertification. — 15th Congress of the R.C.M.N.S. ‘Exploring a physical laboratory: the Mediterranean Basin’, 3—6 September 2017, Athens, Abstracs, p. 31.

Böhme, M., Kötter, S., Fuss, J., Uhlig, G., Spassov, N., 2017. The diet of Graecopithecus freybergi: phytoliths and dentognathic evidence. — 5th Congress of the RCMNS. September 03—06.2017. Book of Abstracts.

Spassov, N., Spiridonov, G., Ivanov, V., Assenov, L., 2016. Bear footprints and their use for monitoring and estimating numbers of brown bears (Ursus arctos L.) in Bulgaria. — Historia naturalis bulgarica, 23: 119—126.

Spassov, N., Ivanov, V., 2016. Home range, movements and activity patterns of an exceptionally large male Brown Bear (Ursus arctos L.) in the area of the Bulgarian-Greek border (Western Rhodope Mts.). — ZooNotes 89: 1—3.

Spassov, N., Ignatov, A., Acosta-Pankov, I., 2016. On the status of the leopard in Turkey, again. — Cat news No 64, Autumn 2016, ISSN 1027-2992.

Spassov, N., 2016. Southeastern Europe as a route for the earliest dispersal of Homo toward Europe: Ecological Conditions and the Timing of the First Human Occupation of Europe. In: Harvati, K., Roksandic, M. (eds), Paleoanthropology of the Balkans and Anatolia. Springer, Dordrecht, 281—290. — 10.1007/978-94-024-0874-4_16

Spassov, N., 2016. On the origin of Wisent, again. — Historia naturalis bulgarica, 23: DOI: 207--209.

Ivanova, S., Gurova, M., Spassov, N., Hristova, L., Tzankov, N., Popov, V., Marinova, E., Makedonska, J., Smith, V., Ottoni, C., Lewis, M., 2016. Magura Cave, Bulgaria: a multidisciplinary study of Late Pleistocene human palaeoenvironment in the Balkans. — Quaternary international, 86—108. DOI: 10.1016/j.quaint.2015.11.082 [PDF]

Hristov, P., Sirakova, D., Mitkov, I., Spassov, N., Radoslavov, G., 2016. Balkan Brachicerous Cattle — the first domesticated cattle in Europe. — Mitochondrial DNA, Part A. DOI: 10.1080/24701394.2016.1238901

Gurova, M., Ivanova, S., Spassov, N., Hristova, L., Krumov, I., Verheyden, S., Marinova, E., Dedov, I., 2016. Excavations at Mishin Kamik Cave: 2015 season. — Archaeological Discoveries and Excavations in 2015, National Archaeological Institute with Museum at the Bulgarian Academy of Sciences, 56—58.

Geraads, D., Spassov, N., 2016. Musteloid carnivores from the upper Miocene of South-Western Bulgaria, and the phylogeny of the Mephitidae. — Geodiversitas, 38 (4): 543—558. DOI: 10.5252/g2016n4a5

David, D., Orr, C. M., Hodgkins, J., Spassov, N., Gurova, M., Miller, C., Tzankov, Ts., 2016. The Human Fossil Record of Bulgaria and the Formulation of Biogeographic Hypotheses. — In: Harvati, K., Roksandic, M. (eds), Paleoanthropology of the Balkans and Anatolia. Springer, Dordrecht, 187—211.

Spassov, N., Tosheva, S., 2015. 125 years National Museum of Natural History, BAS. — Historia naturalis bulgarica, 21: 3—8. [PDF]

Spassov, N., Spiridonov, G., Ivanov, V., Assenov, L., 2015. Signs of the bear life activities and their utilization for the monitoring of the bear (Ursus arctos L.) in Bulgaria. — Historia naturalis bulgarica, 22: 73—83. [PDF]

Spassov, N., Peykov, V., Simeonov, P., 2015. Evidences for the Lynx recovery in Bulgaria: the Lynx discovered in Western Rhodopes. — ZooNotes, 73: 1—4.

Spassov, N., Iliev, N., Karastoyanova, N., Chohadziev, S., 2015. The Remains of Wild and Domestic Animals from the Late Chalcolithic Tell Settlement of Hotnitsa (Northern Bulgaria). — Archaeologia Bulgarica, IXI, 2, 1—21.


— 259 publications
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